Chapter 3.7 Wild plants of primary lowland tropical rain forest

Malcolm Ross

1. Introduction

This chapter focuses principally on the lowland tropical rain forests of New Britain and New Ireland, the large islands of the Bismarck Archipelago, as these are the forests whose margins - at the very least - would have been familiar to Proto Oceanic (POc) speakers. It would be appropriate also to include the rain forests of the Admiralties, but little is known about them (World Wildlife Fund 2007a). Some attention is also given to the other islands of NW Island Melanesia, namely Bougainville and the Solomons, as they were occupied by speakers of early Oceanic dialects which were perhaps barely distinct from POc.

Also included here are trees which grow in freshwater swamp forest, a habitat that is not common in the Bismarcks, though more widespread in Bougainville and parts of the Solomons (ch.2, §3.1.3). Excluded are (i) plants that mainly occur in littoral forest (see chapter 5) and (ii) plants that occur in the wild but are also tended or cultivated to some degree (see chapters 9-11 and 13).

In POc times, the Bismarcks would have been almost completely covered in rain forest, and much of their area remained so until 30 years ago, although there were of course substantial portions of secondary forest resulting from the agricultural activities of Oceanic speakers over 3000 years (ch.2, §3.2).1

Although New Britain and New Ireland have two distinct soil types, one limestone-based, the other volcanic, there is surprisingly little difference in their species composition. Major lowland rain forest tree genera include Pometia, Octomeles, Alstonia, Campnosperma, Canarium, Dracontomelon, Pterocymbium, Crytocarya, Intsia, Ficus, Terminalia and Vitex.

Compared with mainland New Guinea the overall diversity of tree species in Bismarcks lowland forests is not impressive. Missing are the two conifers Araucaria hunsteinii and Araucaria cunninghamii that tower above the lowland broadleaf forests in New Guinea. The dipterocarps (Dipterocarpaceae) that dominate much of peninsular and island southeast Asia are represented by only three species in New Guinea, and probably not at all in the Bismarcks (World Wildlife Fund 2007b).

Freshwater swamp forests on New Britain and New Ireland include Campnosperma brevipetiolata (§4.1), Terminalia brassii, the sago palm (Metroxylon sagu), and species of Pandanus. Limestone forests near the coast of southern New Ireland and along the coast and interior of New Britain are dominated by Vitex cofassus (World Wildlife Fund 2007b).

2. Rain forest layers

A rain forest typically has four or five layers (opinions differ on the division of the lower layers), and these provided the basis for the organisation of this chapter.2

The tallest trees, spaced well apart, jut out above the forest canopy with umbrella-shaped crowns at heights of over 50 m. These emergent trees, treated in §3, have straight, smooth trunks with few branches. They need to be able to withstand high temperatures and drying winds and tend to have small, pointed leaves. Some species lose their leaves during the brief dry season in monsoon rainforests.

The canopy (§4) contains a majority of the larger trees, typically 30–50 m tall, and its adjacent treetops provide a more or less continuous cover of broad-leaved evergreen foliage. The branches of the upper portion of the canopy often support a rich flora of epiphytes, including orchids, mosses, and lichens. The canopy receives plenty of sunlight, but allows only a small amount to penetrate to the layers below. The leaves are formed in such a way as to allow rain to run off. This keeps them dry and prevents mold and mildew from forming in the humid environment. Many canopy tree species have large buttresses at the base of the trunk. Formerly believed to help support the tree, it is now believed that the buttresses channel dissolved nutrients to the shallow root system.

The understorey has two parts: a lower canopy (§5) consisting of trees around 20m in height and a shrub layer (§6) of small trees, shrubs, herbs and ferns able to survive on the 5% of sunlight which reaches the understorey. Understorey plants tend to have large leaves in order to catch as much as possible of the sparsely dappled sunlight: they are largely protected from winds which would damage large leaves in the canopy. Some trees have larger leaves when they are shorter but smaller leaves when they reach canopy height. Large woody vines climb the trees to capture sunlight. There is little air movement and constantly high humidity in the understorey.

The forest floor (§7) receives only 2% of the rainforest’s sunlight, and only specially adapted plants can grow under these conditions. Away from river banks, swamps and clearings, where dense undergrowth is found, the forest floor is relatively empty of vegetation. It also contains decaying plant and animal matter, which decays rapidly in the warmth and humidity to be absorbed by the trees’ shallow roots. Many forms of fungi grow here, assisting in the decay.

3. Emergent trees

Emergent trees are those which regularly grow to 50 m or more. There are also a number of canopy trees that grow taller and acquire emergent status in some localities. These are noted in the various subsections of §4.

3.1. Alstonia scholaris, milky pine, canoe tree, TP kanu (Apocynaceae)

Alstonia scholaris is an emergent or tall canopy tree, growing straight to a height of 30–50 m or more, with few branches on its trunk. It grows only in the forest, often near rivers (Hviding 2005: 146). It is a salient tree in the Bismarcks and the Solomons, but it is rare in Remote Oceania, probably because its use there in constructing large canoes has put the survival of at least taller specimens under pressure (W. McClatchey, pers. comm.).

The straight trunk with its lightweight wood is used in many locations for making canoes, for beams, for shark-trap and fishing-net floats and for carving (Powell 1976, Peekel 1984: 441, Hviding 2005: 146). At Teop (north Bougainville), the tree is known simply as sinivi ‘canoe’ (Record 1945). Wood from buttresses was used for the tall prows of Marovo war canoes.

On New Britain the latex is said to relieve colds (Floyd 1954). At Marovo the sap or bark scrapings boiled in water is a medicine for stomach ache.

At Marovo too, the dead were commonly buried in a sitting position between the buttresses of Alstonia scholaris, and so it was associated with the departure of the spirits of the dead, resulting in taboos against felling it.

A PMP form for ‘Alstonia scholaris’, namely *ditah, is reconstructable (ACD), but I have found no Oceanic reflexes of this. Despite their apparent variation, the reflexes of POc *sabakap below are largely regular. Kara, Patpatar and Nehan reflect regular loss of *-k- but retain the final consonant. NNG languages, Nakanai and SES languages lose the final consonant. The Choiseul languages Varisi, Avaso, Ririo, Babatana and Sisingga reflect Proto Northwest Solomonic *baɣava (for †*abaɣava) in which initial *s- is lost through lenition (a sporadic process) and the final consonant is retained with the addition of an echo vowel.

POc *sabakap Alstonia scholaris’ (Chowning 2001: 84)
NNG Longeinga samvaga Alstonia scholaris’ (Panoff 1972)
NNG Kairiru sabok Alstonia scholaris
MM Nakanai sabaka Alstonia sp.
MM East Kara savəf Alstonia scholaris’ (-f unexplained)
MM Patpatar sabau Alstonia scholaris’ (-u unexplained)
MM Nehan habau Alstonia scholaris’ (-u unexplained)
MM Varisi baɣava Alstonia scholaris’ (W. McClatchey, pers. comm.)
MM Avasö boava Alstonia scholaris’ (W. McClatchey, pers. comm.)
MM Ririo boʔo Alstonia scholaris’ (W. McClatchey, pers. comm.)
MM Babatana bua Alstonia scholaris’ (W. McClatchey, pers. comm.)
MM Sisiqa baɣava Alstonia scholaris’ (W. McClatchey, pers. comm.)
SES Gela habaga Alstonia scholaris
SES Kwaio tabaʔa Alstonia scholaris
SES Kwara’ae tabaʔa Alstonia scholaris
SES Ulawa tabaʔa Alstonia scholaris

3.2. Falcataria moluccana (syn. Albizia falcataria, Albizia moluccana, Albizia falcata, Paraserianthes falcataria) (Mimosaceae)

Figure 7.1: Falcataria moluccana.

Falcataria moluccana was until recently known as Albizia falcataria. Because there are items in the data glossed ‘Albizia sp.’ which may in fact denote Falcataria moluccana, the genera Falcataria and Albizia are handled together here.

The small-leafed crown of Falcataria moluccana emerges above all other trees in Bismarcks rain forests. It grows to between 30 and 60 m in height with a trunk up to 1 m in diameter (Peekel 1984: 207-208), but it is brittle and can come crashing to the ground (W. McClatchey, pers. comm.).

On New Britain and on Manus its trunk is used for canoe hulls, on New Britain also for slitgongs (Powell 1976, Arentz et al. 1989: 94, O’Collins & Lamothe 1989). However, McClatchey points out that it does not serve these purposes well, as objects made from it don’t last. It is more likely to be given to learner carvers or canoe-makers for practice.

The reconstruction of a term (or terms) for Falcataria moluccana is difficult. Kwa’ioloa & Burt (2001: 107), discussing the similar Albizia salomonensis on Malaita, say that it only grows close to rivers and prefers sandy soils. If the same is true of Falcataria moluccana, then its occurrence in Bismarcks rain forests three thousand years ago may have been rather rare, resulting in the frequent loss of inherited terms for the species. Alternatively, the fact that it has little use may account for the dearth of cognates (ch.14, §2).

All three reconstructions below entail uncertainties. POc *babak has just two reflexes. With regard to POc *pail and *kai(k), POc vowel sequences like *-ai- were fairly rare, and it is possible that *q intervened between *a and *i, but *-q- is reflected in none of the modern languages nor in the CMP cognates which support the reconstruction of PCEMP *bail3 and *ka(w)iak.4 The latter is reconstructed with an uncertain medial -w-, reflected in E Sumba kawia[ka]. If -w- was present at an earlier stage, it is irregularly lost in Nakanai kai.

POc *babak Falcataria moluccana
Adm Bipi pap Falcataria moluccana
MM East Kara vavak Falcataria moluccana

PCEMP *bail Falcataria moluccana or Albizia sp.
POc *pail Falcataria moluccana
Adm Nyindrou bei Falcataria moluccana
NNG Lukep pai-pai tree sp. with light wood used for making canoe’s outrigger
MM Tolai vail-ail a beach tree, Pongamia pinnata
MM Nehan puil Falcataria moluccana’ (-ui- for †-ei-)
MM Babatana va-vae (miga) Albizia saman’ (introduced from S America)
MM Kia fai Falcataria moluccana’ (W. McClatchey, pers. comm.)
SES Longgu pai tree sp. used to make canoes
SES Kwara’ae fai Falcataria moluccana
PCEMP *ka(w)iak Albizia sp.
POc *kai(k) Albizia sp.
MM Nakanai kai Albizia sp.’ (Floyd 1954)

3.3. Octomeles sumatrana, TP erima (Datiscaceae)

Octomeles sumatrana is one of the tallest trees in the Bismarck Archipelago and Bougainville, at 40-80 m tall with huge buttresses, above which the trunk of soft white wood is up to 2.5 m in diameter (Peekel 1984: 391). The nectar-rich flowers attract flying foxes (Record 1945). The wood is widely used in the Bismarcks to make canoes (Floyd 1954, Peekel 1984:391, Arentz et al. 1989:93, Floyd 1954, Bugenhagen & Bugenhagen n.d.).

Octomeles sumatrana is apparently of little significance in the Solomons (Whitmore 1966)5 and does not occur at all in Vanuatu, to judge from its absence from Gowers (1976) and Wheatley (1992).

The reconstruction below is for PWOc only, but this is not surprising in view of the limited distribution. Two versions of the reconstruction are supported, *kuRim(a,o) and *iRim(a,o). They overlap geographically, and their initial syllables may simply reflect different prefixes (ch.2, §7.1.2).

Figure 7.2: Octomeles sumatrana.

PWOc *kuRim(a,o), *iRim(a,o) Octomeles sumatrana’ (ACD: *iRimo)
NNG Mangap kurīmi Octomeles sumatrana
NNG Yabem (ka)kelim tree with large leaves and thick, strong trunk’ (identified as Octomeles sp. by Lane-Poole 1925)
PT Motu irimo tree sp. from which canoes are generally made
PT Ubir irim tree sp., used for canoes
PT Tawala ilimo large tree sp., used for war canoes
MM Nakanai (ko)imu Octomeles sumatrana’ (zero for †-l- < *-R-)
MM East Kara ima Octomeles sumatrana
MM Madak ima tree sp.
MM Patpatar irime Octomeles sumatrana
MM Tolai irima Octomeles sumatrana
MM Label irimu Octomeles sumatrana
MM Teop inimo Octomeles sumatrana’ (Record 1945)
MM Babatana vurima Octomeles sumatrana’ (McClatchey et al. 2005)

4. The forest canopy

As a rough rule of thumb, trees of the forest canopy are assumed to be those which grow to between 30 and 50 m. There are also several species, noted in the subsections of §5, which are usually sub-canopy trees but which grow taller in some localities and become part of the canopy itself.

Certain trees of the forest canopy are treated elsewhere in this volume. Strangler figs become part of the forest canopy by using an existing tree as host to piggyback their way into the light (ch.1 0, §4). Canarium species and Terminalia species, especially Terminalia kaernbachii, and Pometia pinnata, are canopy trees in the Bismarcks, but have also long been cultivated, the first two for their nuts and Pometia pinnata for its fruit, and are thus treated in ch.11 (§§2.1, 2.4 and 3.5 respectively).

There are several canopy trees that are only reported from lowland rain forests in the Solomons but not in the Bismarcks, and for some of these no POc term can be reconstructed. This is perhaps significant, given that the POc homeland is believed to have been in the Bismarcks (vol.2, ch.2). These trees are Gmelina moluccana, Pterocymbium species, Schizomeria serrata and Terminalia calamansanai (Mueller-Dombois & Fosberg 1998: 53-54).

4.1. Campnosperma brevipetiolatum (Anacardiaceae)

Campnosperma brevipetiolatum is a large canopy tree, growing up to 50 m in height. Its straight smooth trunk has a cylindrical bole, usually up to 1.2 min diameter and occasionally as much as 2m (Conn & Damas 2006).

The Bola of New Britain use it for canoe hulls (Powell 1976). The Marovo consider it inferior to Gmelina moluccana for this purpose, but good for house planks (Hviding 2005: 134).

Campnosperma brevipetiolatum is apparently not found in Remote Oceania, and its distribution in the Bismarcks and the Solomons seems to be patchy, as it is missing from the usually very thorough Peekel (1984) and Henderson & Hancock (1988). There is just one weakly supported POc term for the species, namely *olaŋa.

POc *olaŋa Campnosperma brevipetiolatum
Adm Bipi laŋ Campnosperma brevipetiolatum’ (O’Collins and Lamothe 1989)
Adm Nyindrou lam Campnosperma brevipetiolatum’ (O’Collins and Lamothe 1989)
MM Marovo olaŋa Campnosperma brevipetiolatum

4.2. Cinnamomum spp., wild cinnamon (Lauraceae)

Trees of the genus Cinnamomum grow to 30 m. They are known for their barks, which are widely processed to make spices and to extract essential oils. Only one of the items in the cognate set supporting POc *(m,mʷ)aso(q)uCinnamomum sp.’ includes a species-level identification within the genus Cinnamomum: Lou moso is glossed as Cinnamomum xanthoneuron, a ‘wild cinnamon’. This is not one of the three species that provide commercial cinnamon bark,6 but one of two tree species exploited by German traders on the north coast of New Guinea under German colonial rule and into the 1930s for their essential oils. The other was not a Cinnamomum species but Cryptocarya aromatica (syn. Cryptocarya massoy, Massoia aromatica), and there was much confusion as to which oil came from the bark of which tree. The oils from these two species are known as lawag oil and massoia oil.7 It is possible that Mager’s gloss of the Bing and Gedaged reflexes as Cryptocarya aromatica reflects this confusion and that the intended denotatum was Cinnamomum xanthoneuron. At any rate, it seems likely that POc *(m,mʷ)aso(q)u did indeed denote Cinnamomum xanthoneuron. The fragrance of Cinnamomum xanthoneuron (and/or Cryptocarya aromatica) bark was known to the traditional residents of north New Guinea and the Bismarcks, as Mager (19 52: 204) reports in his gloss of the Gedaged and Bing items,

The bark is used a great deal in sorcery. It is chewed and spit [sic] out into the face of the spirits, so as to drive them away. A piece of bark is carried in the net bag to keep evil spirits from harming the child.

Arentz et al. (1989: 92) also report that on New Britain the bark is consumed as a medicine against fever and stomach pain.

POc *(m,mʷ)aso(q)u has cognates in the languages of Java: Sundanese maŋsoi, Javanese masoyi, masogi, Madurese masoji, all denoting massoia oil rather than a tree species. The species label ‘massoy’ and the term ‘massoia’ are probably derived from the Javanese term, but, as a result of the confusion between the two oils, are applied to Cryptocarya aromatica, syn. massoy, and its essential oil, rather than to Cinnamomum xanthoneuron. It seems likely, incidentally, that Are masoɣi (for †masou) is a borrowing, perhaps indirectly, from Javanese, rather than a directly inherited reflex of *(m,mʷ)aso(q)u.8

The reflexes of POc *(m,mʷ)aso(q)u reveal a further set of complications. (Blust 1981a) glosses the Lou and Nauna terms ‘Cananga odorata’ (§5.2), rather than a Cinnamomum species, and this is also the gloss of the Meso-Melanesian reflexes. Fijian has two terms: Bauan maðou ‘wild cinnamon, Cinnamomum sp.’ and Bauan makosoi / Wayan mākosoi, both ‘Cananga odorata’. The latter appear to be metathesised forms of a PCP compound *(m,mʷ)aso-koi ‘perfume tree, Cananga odorata’.9 This raises the possibility that POc *(m,mʷ)aso(q)u denoted Cananga odorata as well as cinnamon, i.e. that it denoted a taxon of perfumed trees. PCP then seems to have distinguished between *(m,mʷ)aso ‘cinnamon’ and *(m,mʷ)aso-koiCananga odorata’.

The possibility that there was such a taxon is strengthened by an observation by Will McClatchey (pers. comm.) that the quote from Mager above could also be applied to the Fijian and Western Polynesian use of Cananga odorata.

The Meso-Melanesian forms below, apparently reflecting PMM *mud(e)u (rather than †*moso(u)) are problematic. They may reflect a PMM borrowing or a non-cognate chance resemblance.

POc *(m,mʷ)aso(q)u wild cinnamon, Cinnamomum sp., probably Cinnamomum xanthoneuron; possibly also Cananga odorata’ (Milke 1968)
Adm Mussau mosou wild cinnamon, Cinnamomum sp.
Adm Lou moso tree with redolent bark, the cinnamon, Cinnamomum xanthoneuron
Adm Baluan mʷasow wild cinnamon, Cinnamomum sp.
Adm Nauna moso Cananga odorata’ (Blust 1981a)
NNG Mengen miau Cinnamomum sp.
NNG Kove modou aromatic plant, possibly cinnamon, used in healing’ (A. Chowning, pers. comm.)
NNG Yabem mʋsı wild cinnamon, Cinnamomum sp.
NNG Bing miyou Cryptocarya aromatica’ (Mager 1952: 204)
NNG Gedaged mio Cryptocarya aromatica’ (Mager 1952: 204)
NNG Megiar muyou cinnamon bark’ (Kasprus 1945)
PT Are masoɣi wild cinnamon, Cinnamomum sp.’ (borrowed ?)
MM East Kara mədeu Cananga odorata
MM Nehan mudu-mud Cananga odorata
MM Varisi mudu-mudu Cananga odorata’ (McClatchey et al. 2005)
MM Ririo mud-mud Cananga odorata’ (W. McClatchey, pers. comm.)
MM Babatana mudu-mudu Cananga odorata’ (McClatchey et al. 2005)
MM Nduke mu-mudu Cananga odorata
MM Marovo mudu a tree of the secondary forest, with yellow fragrant flowers that are used in coconut oil
Fij Bauan maðou wild cinnamon, Cinnamomum sp.
PCP *(m,mʷ)aso-koi perfume tree, Cananga odorata’ (Milke 1961)
Fij Wayan mākosoi Cananga odorata’ (metathesis)
Fij Bauan makosoi Cananga odorata’ (metathesis)
PPn *mosokoi Cananga odorata
Pn Tongan mohokoi Cananga odorata
Pn East Futunan mosokoi Cananga odorata
Pn Tikopia mosokoi Cananga odorata
Pn Samoan mosoʔoi Cananga odorata
cf. also:
SES Kwara’ae mudu Dillenia ingens
Fij Rotuman moskoy tree with greenish-yellow flowers and clusters of fruit; timber used for canoes’ (Polynesian borrowing)

4.3. Dillenia schlechteri (syn. Dillenia macrophylla) (Dilleniaceae)

There are many Dillenia species in SE Asia and Oceania, some of them tall canopy trees, others smaller trees of the lower canopy.10 Only one species is reported from the Bismarcks (Peekel 1984: 375, Conn & Damas 2006), Dillenia schlechteri, a canopy tree 30–50 m tall with a light red trunk. However, there is linguistic evidence in the form of POc *drokol (§5.4) that at least one sub-canopy species was known to POc speakers.

In the Admiralties Dillenia schlechteri is used for house construction. The timber is said to last over 30 years if it is not exposed to the elements (O’Collins & Lamothe 1989).

The second and third vowels and the possible final consonant of POc *kulapu(R)Dillenia schlechteri’ are due to the reconstruction of PMP *kelabuR ‘large Dillenia species’ on the basis of the data below and of Blit Manobo klambug (daka)Dillenia megalantha’ (cf§5.4), Bagobo kalambok, Lanao kalambuguy,11 both ‘Dillenia philippinensis’ (Madulid 2001b: 100).

PMP *kelabuR large Dillenia species
POc *kulapu(R) Dillenia schlechteri
Adm Nyindrou kun Dillenia sp.
MM Patpatar (e)kulap Dillenia schlechteri
Fij Bauan kuluva Dillenia biflora’ (Keppel et al. 2005)
Fij Wayan kulu-kulu Dillenia biflora12

4.4. Dracontomelon dao (syn. Dracontomelon mangiferum, Dracontomelon edule), New Guinea walnut, Dragon plum, TP mon, B nakatambol (Anacardiaceae)

Figure 7.3: Dracontomelon dao, New Guinea Walnut.

The New Guinea walnut tree, Dracontomelon dao, is massive, usually reaching 30-35 m, and occasionally 50 m, in height. It has large buttresses and above the buttresses often has a circumference of 3-5 m. At a height of about 7-10 m the trunk divides into a pair of large branches. Each of them continues to divide upwards and sideways recursively to form a large umbrella-shaped crown. The leaves are made up of 6 to 10 leaflets with a smooth edge (Figure 7.3). The fruit are small, 2-3 cm in diameter, and have five flecks around them. The small amount of flesh around the flattened seed is edible but tart and is consumed fresh (Peekel 1984: 323, French 1986: 238). Dracontomelon dao occurs from SE Asia to the Solomons (Walter & Sam 2002: 158).

Bourke (in preparation, n.d.) writes that although Dracontomelon dao was traditionally a significant fruit in the area around Madang, it is unimportant or absent elsewhere in the lowlands of New Guinea. Places where it is recorded as being eaten are the Schouten Islands off the mouth of the Sepik River, some of the small islands in the Admiralties, some islands in SE Papua, the Duke of York Islands (between New Britain and New Ireland), Nissan Island (between New Ireland and Bougainville) and Bougainville. The only report of cultivation comes from French, who says it is sometimes planted from seed.

Two terms are reconstructable: POc *raqu(p) and PNCV *katabola. The final bracketed *-p of *raqu(p) is added to take account of th1e final consonants of the Patpatar and Tolai reflexes. However, there are no known non-Oceanic reflexes of *-p. The PCP *tawa-raquDracontomelon vitiense’ contains a reflex of POc *tawanPometia pinnata’ (ch. 11, §3.5) as its first element, as the fruits of the two plants are similar in appearance (Geraghty 2004: 80).

PAn *daqu Dracontomelon dao’ (Blust 1986)
POc *raqu(p) New Guinea walnut, Dracontomelon dao
Adm Mussau ra Dracontomelon dao
Adm Nyindrou ⁿrau Dracontomelon sp.
Adm Baluan you Dracontomelon dao
NNG Lukep rak Dracontomelon dao
NNG Takia rau Dracontomelon dao
MM Nakanai lahu a tall tree (Anacardiaceae) used for planks
MM Tolai laup Dracontomelon dao’ (for †rau)
MM Patpatar loh Dracontomelon dao’ (for †ro)
PROc *raqu dragon plum tree, Dracontomelon dao
NCV Mwotlap ye Dracontomelon dao
NCV Mota rau a fruit tree
NCV Paamese e-au Dracontomelon dao
NCV Lewo (puru-)lu Dracontomelon dao
NCV Namakir raʔ Dracontomelon dao
NCV Nguna na-rau Dracontomelon dao
NCV Nguna na-rau Dracontomelon dao
SV Sye na-raɣ Dracontomelon dao
SV Kwamera nə-rai Dracontomelon dao
PCP *(tawa)raqu dragon plum tree, Dracontomelon vitiense
Fij Wayan (tawa)rau Dracontomelon vitiense
Fij Rotuman (jav)rau Dracontomelon vitiense’ (Geraghty 2004: 80)
Pn Emae (tava)rau Dracontomelon vitiense’ (Geraghty 2004: 80)
Pn West Futunan (tave)rau Dracontomelon vitiense’ (Geraghty 2004: 80)

PNCV *katabola Dracontomelon dao’ (Clark 1996)
NCV Ambae gatabola Dracontomelon dao
NCV Kiai atapolo Dracontomelon dao
NCV Raga ɣatabola tree sp.’ (Walsh 2004)
NCV Tamambo (vu)hatabola tree sp.
NCV Big Nambas na-hatapul Dracontomelon dao
NCV Uripiv ni-tapol Dracontomelon dao
NCV Naman n-atabal Dracontomelon dao
NCV Neve’ei na-ʔatebʷel Dracontomelon dao
NCV Avava atibol Dracontomelon dao
NCV Nese ɣatabol Dracontomelon dao

4.5. Dysoxylum spp., stinktree, stinkwood, B stingwud (Meliaceae)

Figure 7.4: Dysoxylum arborescens.

A number of Dysoxylum species grow in NW Island Melanesia, ranging from the tall canopy tree Dysoxylum gaudichaudianum (syn. D. amooriodes) sometimes growing to 35 m, to the sub-canopy tree Dysoxylum kaniense. An important tree between these extremes is Dysoxylum arborescens, also a canopy tree, but usually only 20–30 m high (Wheatley 1992: 157-160, Kwa’ioloa & Burt2001: 181, Conn & Damas 2006).

A salient feature of all Dysoxylum species is their strong smell, which varies from species to species: some are unpleasant, some pleasant (W. McClatchey, pers. comm.). The unpleasant smell of certain Dysoxylum species has earned them the English name ‘stinktree’ or ‘stinkwood’: when the bark is stripped off and fresh wood is exposed, Dysoxylum gaudichaudianum is said by the Kwara’ae to smell like a man who has not washed and Dysoxylum kaniense to have a smell that induces vomiting (Kwa’ioloa & Burt 2001: 122, 181). Whistler (1991b: 93) comments that the leaves have ‘a disagreeable odor’. Despite the smell, the wood of Dysoxylum species is a useful hardwood, and the timber of the larger species is widely used for house posts (Floyd 1954, Whistler 1991b: 93, Wheatley 1992: 157, 162, Kwa’ioloa & Burt 2001: 122).

POc *maqota perhaps denoted a taxon including several Dysoxylum species. All the species mentioned in the glosses below are tall canopy trees except Dysoxylum kaniense and Dysoxylum spectabile, but this is perhaps because the larger species are more widespread and more salient.

Lynch (2001c: 242) attributes the SV members of the two cognate sets below to a single PSV etymon *ni-m(d,t)awan, but they appear to reflect two etyma. PSV *nə-mtaw reflects metathesis of a variant form *mawota, also reflected in Bauan Fijian mavota. Proto Erakor-Tafea (Lynch 2001c: 189) *tuan is a separate etymon.13

POc *maqota Dysoxylum spp.
PT Muyuw (a)mʷakot a 20-metre Dysoxylum sp.
SES Kwara’ae maoa Dysoxylum kaniense
NCV Mwotlap ma-mot Dysoxylum arborescens
NCV Mota ma-maota tree sp.
PSV *na-mtaw Dysoxylum sp.’ (Lynch 2001c)
SV Sye ni-mtu Dysoxylum aneityense
SV Anejom̃ ne-mtav Dysoxylum aneityense
Fij Bauan mavota Myristica grandiflora’ (for †maota)
Pn Tongan moʔota taxon of three Dysoxylum spp., but primarily Dysoxylum forsteri’ (’Whistler 1991b: 92-93)
Pn Niuean moota tree, Dysoxylum richii, timber used to build main hull of canoe’ (also maota)
Pn East Uvean maʔota Dysoxylum samoensis
Pn East Futunan māʔota shrub sp., Dysoxylum forsteri
Pn Rennellese maʔota tree sp., Dysoxylum gaudichaudianum, valuable for house timbers
Pn Samoan maota Dysoxylum spp.’ (Whistler 2000: 181)
Pn Takuu maota hard, red, wood that drifts to Takuu
Pn Māori māota Dysoxylum spectabile
Proto Erakor-Tafea *tuan Dysoxylum spp.
NCV South Efate ne-tue Anthocarapa nitidula’ (Wheatley 1992: 157)
SV Lenakel ne-tuan Dysoxylum gaudichaudianum
SV Kwamera nə-tuan Dysoxylum gaudichaudianum

4.6. Elaeocarpus spp. (Elaeocarpaceae)

Elaeocarpus species in NW Island Melanesia include the large canopy trees Elaeocarpus angustifolius (syn. Elaeocarpus sphaericus) with its distinctively cornflower-blue fruit and Elaeocarpus floridanus, more common in the Solomons than the Bismarcks, with dark blue or black fruit (syn. Elaeocarpus pseudosepicanus), both 25 or more metres tall, as well as shrub-sized trees of the understorey like Elaeocarpus edulis (syn. Aceratium oppositifolium), 3-6 m tall, with its 4 x 3cm ellipsoid apple-red fruit. The fruits of all species are inedible (Peekel 1984: 352-353, Wheatley 1992: 85, Kwa’ioloa & Burt 2001: 123).

The large species provide wood for house beams at various locations (Record 1945, Arentz et al. 1989: 94), but Will McClatchey (pers. comm.) finds that it is not useful in the traditional communities which he has studied.

Although plenty of names for Elaeocarpus species have been collected, surprisingly few of them form cognate sets - none at all for POc or Western Oceanic, and one each for PEOc and PSV. The absence of a POc or PWOc term perhaps exemplifies the principle that the names of less useful plants are easily forgotten (§2), so that present-day names reflect new coinages at different island Melanesian localities. Alternatively, it may be that Elaeocarpus species were rare in the environments of early Oceanic speakers: the botanical literature suggests that large Elaeocarpus species are significant contributors to the canopy of lowland rain forests in Bougainville and the Solomons but not in the Bismarcks where POc was spoken (Paijmans 1976:64–65, Mueller-Dombois & Fosberg 1998:53-54, 60–61).

Two reconstructions are presented below. PEOc *meloElaeocarpus angustifolius’ is supported by just two reflexes, but the languages are sufficiently far apart to preclude borrowing.

PEOc *melo Elaeocarpus angustifolius
SES Kwara’ae melo Elaeocarpus angustifolius
NCV Vera’a mel Elaeocarpus angustifolius
PSV *na-(s,j)u(v,w)as Elaeocarpus angustifolius’ (Lynch 2004a)
SV Sye ne-yoh Elaeocarpus angustifolius
SV Kwamera nə-suvas k.o. tree with edible seeds in a hairy pod
SV Anejom̃ na-woθ Elaeocarpus angustifolius

4.7. Endospermum spp., cheesewood, milkwood, whitewood, PNG basswood, B waetwud, melektri (Euphorbiaceae)

Within island Melanesia Endospermum medullosum (Figure 7.5, left) is present in the Bismarcks, the Solomons and Vanuatu; Endospermum moluccanum is reported only from the Bismarcks (Peekel 1984: 313, Wheatley 1992:91, Hviding 2005: 135). Occasionally an emergent tree, otherwise a large canopy tree up to 45 m high, Endospermum medullosum has a markedly fluted bole of 45-100 cm diameter, sometimes crooked, sometimes straight and up to 25 m long. It has a distinctive crown, shallow, flat-topped and umbrella-like. Its distribution is limited by the fact that it does not tolerate shade: it grows well only where there are gaps in the canopy (Thomson 2006a).

Figure 7.5: Left Endospermum medullosum, cheesewood,: A, tree; B, leaves and flowers; C, fruit. Right Pterocarpus indicus, New Guinea rosewood: A, tree; B, portion ofleafwith leaflets; C, shoot with pods.

A somewhat smaller but similar member of the genus is Endospermum moluccanum (syn. Endospermum formicarum),14 sometimes a canopy tree up to 25m high, sometimes a sub-canopy tree (Conn & Damas 2006).

Both species have lightweight wood that is used for fishing-net floats. Peekel (1984: 315) reports from New Ireland that the pith of the twigs of Endospermum moluccanum is usually destroyed by black ants, which inhabit the resulting space. Kwa’ioloa & Burt (2001: 115) report from Kwara’ae that Endospermum medullosum is not used in house construction because the wood is eaten by insects.

POc *koma(r,R)(o,u) may have referred to either or both species. Like the large Elaeocarpus species (§4.6), Endospermum species are not prominent in Bismarcks rain forests, and appear to have undergone a good deal of local re-naming.

POc *koma(r,R)(o,u) Endospermum sp.
MM Nakanai ko-komalu Endospermum moluccanum
NCV Mwotlap no-komʷa Endospermum sp.
NCV Araki (vi)kunᫀaro Endospermum medullosum
NCV Tangoa (vi)kumaro Endospermum medullosum’ (Gowers 1976: 73)
NCV Sakao (du)gomara Endospermum medullosum’ (Gowers 1976: 73)

4.8. Garuga floribunda (syn. Garuga abilo), B namalaus (Burseraceae)

Garuga floribunda is a canopy tree which grows to a height of up to 25m in Vanuatu and up to 35m in Papua New Guinea (Wheatley 1992:63, Conn & Damas 2006). It has a short bole and steeply rising branches, often with a flat crown (Johns 1976: 195). According to Paijmans (1976: 52) its occurrence is limited to the few locations where there is a marked dry season.

There is apparently a traditional perception that Garuga floribunda resembles the smaller Spondias cytherea, noted by Peekel (1984: 283) for New Ireland and reflected in PNCV *mala-usiGaruga floribunda’, reconstructed below. The perception is apparently based on the fact that both trees lose their leaves and are bare between flowering and fruiting. PNCV *mala-usi consists of a reflex of the POc prefix *mala- ‘resembling’ (ch.2, §7.1.4) plus PNCV *usiSpondias cytherea’ (ch.ll, §3.6), i.e. its original meaning was evidently ‘resembing Spondias cytherea’. As noted below, there are also unprefixed reflexes of PNCV *usi which denote Garuga floribunda.

PCP *manauiGaruga floribunda’, also below, appears cognate with the PNCV form, but the medial consonants do not correspond, suggesting that one of the two forms is the result of borrowing. Since PNCV *mala- regularly reflects POc *mala-, but PCP *mana- (instead of expected †*mala-) does not, it is the Central Pacific forms that appear to reflect a borrowing.

PNCV *malausi Garuga floribunda’ (from data in Wheatley 1992)
NCV Mota mʷa-mʷalau Garuga floribunda
NCV Ambae malawhi Garuga floribunda
NCV Nduindui malaouk Garuga floribunda
NCV Tamambo (vu)malaus Garuga floribunda
NCV Tangoa (vi)malaus Garuga floribunda
NCV Tolomako na-malaus Garuga floribunda
PCP *manaui Garuga floribunda
Fij Wayan manawī a tree of dry forest, Rhus taitensis15
Pn Tongan manaui Garuga floribunda
Pn East Futunan manaui large forest tree, Myristica hypargyraea
Pn Samoan maŋaui a large tree, Garuga floribunda’ (Whistler 2000: 179)
PNCV *usi mummy apple, Spondias cytherea’ (from data in Wheatley 1992)
NCV Maewo o-us Garuga floribunda
NCV Nduindui uhi Garuga floribunda
NCV Tolomako na-us Garuga floribunda
NCV Tangoa (vi)usi Garuga floribunda
NCV Uripiv na-us Garuga floribunda

4.9. Intsia bijuga (syn. Intsia amboinensis, Afzelia bijuga), ironwood, island teak, TP kwila, B natora (Cisalpiniaceae)

One of the larger trees of the forests of NW Island Melanesia, Intsia bijuga grows 40-45 m tall and sometimes more (Figure 7.6, left). It is common on the foreshore, but also occurs in lowland rain forests (Peekel 1984: 214-216).

Intsia bijuga is considered one of the strongest and most durable woods in NW Island Melanesia and Vanuatu. It seasons slowly with very little shrinkage and is durable in the ground, resistant to termites and moderately durable in salt water (Gowers 1976: 91). Its uses range from house posts and floorboards to axe handles, slitgongs and wooden bowls (Streicher 1982, Arentz et al. 1989:94, Whistler 1991b: 125, Hviding 2005: 122, F. Damon, pers. comm.). Damon reports that on Woodlark Island it was used for the steering oars of the large canoes that plied the eastern half of the Kula ring.

Superficially, it looks as if there were two POc terms for Intsia bijuga: *toRas and *qipil. However, Blust’s gloss of PMP *teRas as ‘hard, hardwood’ suggests that POc *toRas too may have denoted ‘hardwood’ in general or a taxon of hardwood trees, rather than Intsia bijuga in particular. This would explain why some of its reflexes denote other hardwood trees. In particular, its Proto Polynesian reflex, *toa, was reapplied to Casuarina equisetifolia. In Muyuw the term meikʷ is used both for Intsia bijuga and for the hard heartwood of any tree (F. Damon, pers. comm.). Thus one word for hardwood has been replaced by another in naming Intsia bijuga.

It seems that PMP *teRas / POc *toRas was, in one of its senses, a stative verb meaning ‘hard, durable’. PMic had a stative verb *ma-toa ‘be firm, hard strong’ (Bender et al. 2003: 54), and tree-denoting reflexes sometimes occur with verbal morphology. In CMP languages we find W Sumba, E Sumba kandora, E Sumba mandoraCalophyllum inophyllum’ (Verheijen 1990: 197), reflecting the PMP stative prefixes *ka- and *ma- respectively (Evans & Ross 2001).

Polynesian languages, meanwhile, have reapplied the PCP reflex of POc *pesi ‘a coastal forest tree, perhaps Pongamia pinnata’ to Intsia bijuga (ch.5, §5.12).

PMP *teRas hard; hardwood’ (Blust 1972a)
POc *toRas a taxon of hardwood trees including Intsia bijuga’ (?)
Adm Seimat tor Intsia bijuga’ (-r for †: borrowed?) (Sorensen 1950)
Adm Lou to Intsia bijuga
NNG Kairiru tor Intsia bijuga
MM Sursurunga toraha a hardwood tree
MM Nehan toraha tree sp. with strong yellowish-white wood, used for carving slitgong drums and paddles
MM Halia tolasa same tree: sp. as Nehan toraha
MM Petats tolas Vitex monophylla
MM Teop tora Vitex monophylla
MM Maringe tʰola type of large softwood tree; canoe with two upraised ends
PEOc *toRa(s) a hardwood tree, Intsia bijuga
SES Lau ola a canoe built of planks sewn together
SES Kwara’ae uʔula Intsia bijuga’ (*o > u: irregular change)
SES Arosi ora tree sp. from which best canoes are made; plank-built canoe
NCV Mwotlap no-toy Decaspermum neoebudicum
NCV Merlav tor Casuarina equisetifolia’ (François 2004b)
NCV South Efate na-tor Intsia bijuga
NCV Mota tora a timber tree
NCV Ambae tora Intsia bijuga
NCV Tamambo (vu)tora Intsia bijuga
NCV Raga tora Intsia bijuga
NCV Atchin tor tree sp., used for canoes, posts, etc.
NCV Nese na-toɾ Intsia bijuga
NCV Uripiv na-toɾ Intsia bijuga
NCV Paamese a-to tree sp. with sap which stings
Fij Bauan doa the heartwood of a tree, solid and dark
PPn *toa Casuarina equisetifolia’ (POLLEX)
Pn Niuean toa Casuarina equisetifolia
Pn Tongan toa Casuarina equisetifolia
Pn East Uvean toa Casuarina equisetifolia
Pn East Futunan toa Casuarina equisetifolia
Pn Rennellese toa Casuarina equisetifolia
Pn Emae toa Casuarina equisetifolia
Pn Tikopia toa Casuarina equisetifolia
Pn West Futunan toa Casuarina equisetifolia
Pn Ifira-Mele toa Casuarina equisetifolia
Pn Samoan toa Casuarina equisetifolia
Pn Tahitian toa Casuarina equisetifolia
Pn Tuamotuan toa Casuarina equisetifolia
Pn Marquesan toa Casuarina equisetifolia
Pn Māori toa-toa Phyllocladus glaucus

The inherited meaning of POc *qipil was apparently ‘Intsia bijuga’, but the glosses of its reflexes suggest that it also denoted ‘Casuarina equisetifolia’. Note that ‘ironwood’ and ‘kwila’ both refer to ‘Intsia bijuga’.

PMP *qipil a hardwood tree, Intsia bijuga’ (ACD; Dempwolff 1938: *ipil)
POc *qipil a taxon of hardwood trees including Intsia bijuga and Casuarina equisetifolia’ (ACD: *(q)ipil)
Adm Drehet ʔih ironwood
Adm Likum ih Casuarina equisetifolia
Adm Nyindrou eih ironwood
MM Tolai ip tree sp.
MM Teop ivin a hardwood tree, kwila
MM Mono-Alu ihili Intsia bijuga’ (W. McClatchey, pers. comm.)
MM Ririo kivil Intsia bijuga
MM Babatana kivili Casuarina equisetifolia
MM Marovo kivili Casuarina equisetifolia
MM Roviana kifli Intsia bijuga’ (W. McClatchey, pers. comm.)
MM Kia ivili Intsia bijuga
MM Maringe khifli Intsia bijuga’ (W. McClatchey, pers. comm.)
NCV Mwotlap n-ip Casuarina equisetifolia
Mic Kiribati ibi tree like Calophyllum inophyllum, but harder and heavier
Pn Samoan ifi(fatu) Intsia bijuga
Pn Samoan ifi(lele) a hard-grained Intsia bijuga

Figure 7.6: Left Intsia bijuga, ironwood: A: young tree; B, leaves and flowers; C, seed pod. Right Vitex cofassus, New Guinea teak: A, tree; B, flowering stem with leaves and portion of terminal inflorescence; C, portion of fruiting inflorescence and stem.
PWOc *bʷana Intsia bijuga
NNG Manam bʷana Intsia bijuga
NNG Yabem (ka)bʷɛŋ ironwood
MM Bola bana ironwood
MM Nakanai bala Intsia bijuga

4.10. Planchonella spp., B komtri (Sapotaceae)

Planchonella species range in size from large canopy trees to small trees of the sub-canopy. The species division of the genus Planchonella remains controversial (Walter & Sam 2002: 226), partly, it seems, because there is considerable variation within and across species. This is perhaps due to past domestication, but the literature gives little indication of present-day cultivation.16 Borrell (1989: 134) identifies six species on Kairiru Island, two of which he is unable to name. There is also an overlap with the genus Pouteria, in the sense that species that have at one time or another been placed in the genus Planchonella have at others been placed in the genus Pouteria. Walter & Sam (2002: 226–227) include Planchonella grayana in their catalogue of fruit trees but say that it is little consumed (because the pulp irritates the gums) except at Tasmate (west Santo), where the mature fruit is either roasted whole or peeled, then washed to remove latex before it is eaten.

Peekel (1984: 429-431) describes Planchonella peekelii (syn. Sideroxylon peekelii), a tree about 15m tall with small ovoid fruit, but he does not mention consumption or any other use.

Depending on location, Planchonella costata (syn. Sideroxylon costa tum) varies in the Solomons and Vanuatu between a small sub-canopy tree and a large tree of the canopy. Its bark ranges from pale grey to black-brown, and it has long narrow leaves and round fruit 3-4cm in diameter. It has a preference for the beach or inland for moist habitats. Its wood is close-grained and can be finely worked: combs can be made from a single flat piece of wood (Gowers 1976: 113, Kwa’ioloa & Burt 2001: 140–141). Wheatley (1992) identifies Planchonella grayana as a variety of Planchonella costata. Gowers (1976: 115) also describes Planchonella linggensis, labelled the ‘comb tree’, leading one to wonder if this also is a variety of Planchonella costata.

In view of this variation we can be sure only that POc *kalaka denoted a species of Planchonella, and perhaps several. Kairiru lalak may reflect a reduplication of suffixless *laka after deletion of the (apparent) prefix *ka- ‘tree’ (ch.2, §7.1.2).

POc *kalaka Planchonella sp.
NNG Kairiru lalak Planchonella obovoidea
MM Tolai kalakala tree sp.
NCV Raga ɣaraŋa Planchonella sp.
NCV Uripiv na-klak Planchonella/ Pouteria spp.
PCP *kalaka Planchonella sp.
Fij Bauan galaka Planchonella costata
Pn Niuean kalaka Planchonella sp.
Pn Tongan kalaka Planchonella samoensis
Pn East Futunan kalaka tree sp.
Pn Emae kalaka tree sp.
Pn Samoan alaʔa Planchonella garberi
Pn Hawaiian ʔālaʔa Planchonella sandwicensis
Pn Tuamotuan karaka tree sp.
Pn Māori karaka Corynocarpus laevigata

The glosses in the cognate set below imply that PCP *bau had the meaning which it retains in Wayan Fijian: ‘Tree … taxon: generic, includes species of Burckella, Manilkara, Palaquium and Planchonella (Sapotaceae)’ (Pawley & Sayaba 2003; see also ch.3, §3 .1). They are all medium to large rain forest trees of the family Sapotaceae. I had hesitated to assume that Teop bauLeea tetramera’ is cognate, as Leea tetramera is a shrub, but Will Mc-Clatchey (pers. comm.) argues that all the plants listed below except Guettarda speciosa are hardwoods from which a useful club could be made and that this fact is central to the definition of the taxon. This being so, it is possible that POc *bau is identical with POc *bouFagraea spp.’ (ch.5, §5.6).

POc *bau hardwood taxon’ (see above)
MM Teop bau Leea tetramera
PCP *bau hardwood taxon’ (see above)
Fij Wayan bau woody trees taxon including Burckella richii, Manilkara vitiensis, Palaquium fidjiense and Planchonella species
Fij Bauan bau Palaquium spp. (Keppel et al. 2005)
Fij Yasawa bau tree sp., probably Sapotaceae sp.
Pn Anutan pau Pipturus argenteus
Pn Tuvalu pau Guettarda speciosa, Mammea glauca
Pn Rennellese pau Planchonella sp.
Pn Samoan pau tree sp. Manilkara hoshinoi, from which clubs are made

4.11. Pterocarpus indicus, New Guinea rosewood, flame wood, B nananara, navilae, bluwota (Fabaceae)

In the Bismarck Archipelago Pterocarpus indicus grows in lowland rain forests, but in the Solomons it is more salient in freshwater swamp forests (Figure 7.5, right). It grows to varying sizes depending on its immediate environment. Sometimes it is an emergent tree up to 40 m high, often it is a large canopy tree, and under some conditions a tree of the lower canopy only 10m high. Like most canopy trees, it has a long bole (which is sometimes crooked) and buttresses which sometimes extend into flutes up the bole (Conn & Damas 2006, Henderson & Hancock 1988: 165, 320).

The flowers are small but bright yellow and fragrant. Its fruit comes in the form of disk-shaped pods. It has red sap, and one of its English names, ‘flame wood’, reflects the fact that its wood is multi-coloured: yellow, red and brown. Its alternative Bislama name bluwota reflects the fact that the wood and bark turn steel blue when immersed in water (Wheatley 1992: 145).

Pterocarpus indicus provides excellent timber, moderately soft, moderately light and permanently sweet-smelling. In various parts of the Solomons and Vanuatu the trunks are used for dugout canoes and planks and for carving (Henderson & Hancock 1988: 167, Wheatley 1992: 145, Kwa’ioloa & Burt 2001: 119, Hviding 2005: 142). It has similar uses in the Bismarcks, where it is also used for hourglass drums (Floyd 1954, Powell 1976).

In a variety of locations from the Bismarcks to Vanuatu an infusion of the young leaves or of the bark is used against diarrhoea or against excessive menstruation (Henderson & Hancock 1988: 167, Arentz et al. 1989: 92, Bourdy & Walter 1994).

Two POc terms for Pterocarpus indicus are reconstructable: *naRa and *Rigi.

PMP *naRa Pterocarpus indicus’ (Blust 1980b; ACD)
POc *naRa Pterocarpus indicus
Adm Lou na Pterocarpus indicus’ (Holdsworth and Wamoi 1981)
Adm Baluan nay tree with red wood, probably Pterocarpus indicus
NNG Gitua nara Pterocarpus indicus
NNG Tami nal Pterocarpus indicus
PT Motu nara Pterocarpus indicus
MM Bola nara-nara Pterocarpus indicus
MM East Kara naɣa Pterocarpus indicus
NCV Mota na-nara Pterocarpus indicus
NCV Vera’a na-nar Pterocarpus indicus
NCV Raga na-nara Pterocarpus indicus
POc *Rigi rosewood, Pterocarpus indicus’ (Geraghty 1990: PEOc *rike)
NNG Atui (ki)riŋ Pterocarpus indicus’ (Arentz 1989:92)
MM Nduke rigi Pterocarpus indicus
MM Roviana rigi tree sp. which yields a good red timber
MM Marovo rigi Pterocarpus indicus
MM Kia grigi Pterocarpus indicus’ (W. McClatchey, pers. comm.)
MM Maringe grigi Pterocarpus indicus’ (Henderson and Hancock 1988)
SES Bugotu ligi Pterocarpus indicus’ (W. McClatchey, pers. comm.)
SES Gela ligi tree sp.
SES Bauro riki Pterocarpus indicus
SES Kwara’ae liki Pterocarpus indicus
SES Lau liki Pterocarpus indicus
SES Arosi rigi Pterocarpus indicus
NCV Bieria (mi)like(he) Pterocarpus indicus
Pn Rennellese li[kq]e tree sp.; barkcloth mallet

4.12. Vitex cofassus, New Guinea teak, TP garamut (Lamiaceae)

Vitex cofassus grows 20-40 m tall (Figure 7.6, right). It has a grey trunk, with a bole up to 18 m long and buttresses up to 6 m high. The wood is smooth, white and durable (Peekel 1984:480, Conn & Damas 2006). At least, this is how specimens of Vitex cofassus in the Bismarcks are described. For the Solomons Henderson & Hancock (1988: 188) describe it as a ‘large, ill-formed tree’, apparently because its buttresses often extend up the bole as irregular flanges and flutings.

A virtue of the hard wood is that it is not eaten by termites (Blewett & Blewett n.d.). In the Bismarck Archipelago it is used for tool handles, and in both the Bismarcks and the Solomons as timber for wall planking, house posts, canoe paddles and canoes, drums and for carving (Powell 1976, Henderson & Hancock 1988: 190, Hviding 2005: 150, Scales n.d.).

The POc term for Vitex cofassus was *pasa(r,R).17 Milke (1961: 171) reconstructed an unglossed POc *pasa on the basis of the Arosi, Sa’a, Bauan Fijian and Samoan items below, together with Gedaged safaCerbera manghas’. He assumes that the latter represents a metathesis, but, given the difference between the two species, this is an ad hoc assumption, and the Gedaged item is omitted here.

Riwo and Takia reflect Proto Bel *payaRi. The presence of the final consonant and added -i points to a loan. Babatana vadaka is also evidently a loan from an unknown source.

POc *pasa(r,R) Vitex cofassus’ (Milke 1961: *pasa)
NNG Manam oara Vitex cofassus
NNG Yalu (a)fas Vitex cofassus’ (Lane-Poole 1925)
MM Nakanai vasa Vitex cofassus
MM East Kara fasei Vitex cofassus
MM Lihir pacere Vitex cofassus’ (Burley 2006)
MM Madak pasa Vitex cofassus
MM Patpatar vasara Vitex cofassus
MM Tolai vara Vitex monophylla
MM Mono-Alu hasala Vitex cofassus’ (Record 1945)
MM Nduke vasara Vitex cofassus
MM Marovo vasara Vitex cofassus
MM Kia varaha Vitex cofassus’ (metathesis; W. McClatchey, pers. comm.)
MM Maringe vahara Vitex cofassus’ (Henderson and Hancock 1988)
SES Bugotu vaha Vitex cofassus’ (W. McClatchey, pers. comm.)
SES Gela vaha Vitex cofassus’ (W. McClatchey, pers. comm.)
SES Lengo vaða Vitex cofassus’ (Henderson and Hancock 1988)
SES Longgu vata Vitex cofassus
SES Lau fata Vitex cofassus
SES Kwara’ae fata Vitex cofassus
SES Arosi hata a large tree sp.
SES Sa’a hata hardwood tree
Fij Bauan vasa Cerbera odollam
Pn Samoan fasa a pandanus species, the leaves of which are used to make mats’ (W. McClatchey, pers. comm.); ‘Pandanus textorius’ (Whistler 2000: 163)
cf. also:
NNG Riwo paiaɬi tree with very hard and durable wood’(Mager 1952)
NNG Takia peari tree with very hard and durable wood’(Mager 1952)
MM Babatana vadaka Vitex cofassus’(McClatchey et al. 2005)

5. The lower canopy

Trees of the lower canopy are assumed to be those with a usual height somewhere between 15 m and 20 m. Again some species display significant height variations and, as noted in the subsections below, are canopy trees in some localities. Conversely, of course, some of the trees described in §4 as canopy trees grow less well in some places and there belong to the lower canopy.

Certain sub-canopy species are treated in other chapters. The barringtonias Barringtonia asiatica, Barringtonia novae-hiberniae and Barringtonia procera are handled in ch.5, §5.2 because of the strong tendency for them to grow on the coast. They are an important part of the lower canopy of the rain forest in parts of Bougainville, however. Other sub-canopy plants, especially in parts of Bougainville and the Solomons, are the betelnut palm, Areca catechu (ch.13, §2.2.1), tree ferns of the genus Cyathea (ch.10, §3.1), fruit trees of the genera Pandanus and Syzygium (ch.ll, §2.5 and §3 .7) (Paijmans 1976:64–65, Mueller-Dombois & Fosberg 1998: 60–61).

No reconstruction could be made for any of the species of Celtis, which are lower canopy trees in the Solomons (Kwa’ioloa & Burt 2001: 157-158), nor for Gnetum latifolium, a large woody climber of Bougainville and the Solomons.

5.1. Bischofia javanica, Java cedar, B nakoka, redwud (Phyllanthaceae)

Occurring throughout the Pacific, Bischofia javanica (Figure 7.7, left) is often a 30–40 m canopy tree with a bole a metre in diameter in the New Guinea region, but is usually a smaller, sub-canopy tree in Vanuatu. Its wood varies from red with red sap to pink or cream with colourless sap. It has light yellowish green flowers and red or black ovoid fruit half a centimetre in diameter, each with six small seeds (Johns 1976:223, Wheatley 1992: 55).

Its occurrence is evidently very patchy. The fact that names for Bischofia javanica have been collected in NW Island Melanesia from Nakanai, Tolai, Roviana and Lau indicates that this species is also found in the locations of these languages, and this is confirmed by Conn & Damas (2006), yet Peekel (1984), Henderson & Hancock (1988), Kwa’ioloa & Burt (2001) and Hviding (2005) make no mention of it, suggesting that in some places it is far from abundant.

Will McClatchey (pers. comm.) offers an explanation of the distribution of Bischofia javanica, commenting that ‘It seems to be in places where they traditionally make clothing from felted bark and not in places where the clothing is made from other sorts of materials. It is of course one of the principal dyes for felted Broussonetia.’

The wood is hard and moderately durable and is used in Vanuatu and Tonga for ground posts (Whistler 1991b: 58, Wheatley 1992: 55)

The PEOc term *koka is reconstructable. The lack of a POc term may simply be due to its absence from a number of sources listing Western Oceanic plant names, i.e. to its odd distribution, rather than to an absence from the environment of POc speakers.

Figure 7.7: Left Bischofia javanica, Java cedar: A, tree; B, compound leaf; C, branch bearing cluster of fruit. Right Cananga odorata, ylang-ylang: A, tree; B, leaves; C, flowering shoot; D, fruit cluster.
PEOc *koka tree sp., Bischofia javanica’ (POLLEX)
SES Ulawa ʔoʔa Bischofia javanica’ (W. McClatchey, pers. comm.)
NCV Tangoa (vi)ɣauha Bischofia javanica
NCV Raga (i)ɣoɣa Bischofia javanica’ (Walsh 2004)
NCV Lewo (puru)koa Bischofia javanica
NCV Nguna na-koka tree sp.
NCV Namakir na-koka Bischofia javanica
NCV South Efate n-kok Bischofia javanica
PCP *koka tree sp., Bischofia javanica
Fij Wayan koka Bischofia javanica
Fij Bauan koka Bischofia javanica
Pn Tongan koka Bischofia javanica
Pn Niuean koka small tree, Baccaurea seemannii
Pn East Futunan koka Bischofia javanica
Pn East Uvean koka Bischofia javanica
Pn Ifira-Mele koka Bischofia javanica
Pn Rarotongan koka Bischofia javanica
Pn Māori koka some edible plant

5.2. Cananga odorata, ylang-ylang, perlume tree, nandingori, nadingro, nangungara (Annonaceae)

Cananga odorata (Figure 7.7, right) is usually a lower canopy tree 10–15 metres tall, but sometimes it reaches the canopy at 30 or 35m (Peekel 1984: 183, Conn & Damas 2006).

Cananga odorata is known across NW Island Melanesia (and elsewhere) for its fragrant yellow flowers, which are dried and used to scent coconut oil (Record 1945, Henderson & Hancock 1988:244, Hviding 2005: 130).

In northern Vanuatu the trunk is hollowed out and used for making drums and the branches are used for outriggers (Gowers 1976:43, Jauncey (in progress)). Kwa’ioloa & Burt (2001: 119) and Wheatley (1992: 42), on the other hand, say that the wood is too soft to be durable and is useful only for internal construction in places with a smoky atmosphere.

Use of the bark and flowers for medicinal purposes is widespread (Floyd 1954, Hviding 2005: 130).

POc *(m,mʷ)aso(q)u may have denoted a taxon which included Cananga odorata (§4.2). POc *pʷi(r,R)a and PSOc *diŋori(q) both appear to have denoted Cananga odorata alone.

POc *pʷi(r,R)a Cananga odorata
MM Lihir pir Cananga odorata
MM Petats bina Cananga odorata
MM Teop bina Cananga odorata
Mic Ponapean pʷur Cananga odorata
Mic Mokilese pʷur tree species
PSOc *diŋori(q) Cananga odorata’ (Clark 1996; Lynch 2004a)
NCV Mwotlap (na-tweh) dıŋıy Cananga odorata’ (François 2004b)
NCV Dorig (wa)dŋʋr Cananga odorata’ (François 2004b)
NCV Ambae diŋori Cananga odorata
NCV Tangoa (ve)riŋori Cananga odorata
NCV Kiai kinori Cananga odorata
NCV Tamambo (vu)diŋori Cananga odorata
NCV Raga diŋori Cananga odorata
NCV Paamese a-reŋe Cananga odorata
SV Kwamera nu-rəŋri tree, sp. wood used for pierced ear and septum ornaments

5.3. Cryptocarya sp. (Lauraceae)

The genus Cryptocarya consists of laurel-like evergreen plants which range in size from lower canopy shrubs to giant emergent trees (Conn & Damas 2006).

Among the large trees are Cryptocarya aromatica, which was exploited by German traders on the north coast of New Guinea for the essential oil in its bark: see §4.2 for discussion. It grows to a height of 45 m and its bole is as much as 1.2 m in diameter. The bark varies in colour, but is often red-brown and corky, with a strong resinous smell. It bears globular berries 10cm in diameter (Johns 1976: 65). Cryptocarya cordata is a similarly large tree, and Record (1945) notes that its currant-sized fruit are used in food as a relish .

The sources of the data supporting POc *nipusCryptocarya sp.’, O’Collins & Lamothe (1989) and Damon (2004), do not provide enough information to allow a species identification, but O’Collins and Lamothe mention that the timber is used in house-building on Manus Island, implying that the species is not too small.18 The sources of the data supporting PWOc *ka(m,mʷ)apaR tell us that the Muyuw tree is more than 10 m tall, the Patpatar tree, dubbed Cryptocarya kamahar by Peekel, 20–30 m tall.

Damon (In preparation) mentions that the nuts have a rich cinnamon-like smell and are important for medicinal purposes on Woodlark Island. Curiously, Muyuw people do not know the tree, as it grows away from areas they frequent, but they know its nuts because birds swallow them whole, then excrete them in their nests, complete with the endocarp, whence they are collected.

Figure 7.8: Cryptocarya aromatica.
POc *nipus Cryptocarya sp.
Adm Bipi ñeu Cryptocarya sp.
Adm Nyindrou nip Cryptocarya sp.
PT Muyuw ni-niwous Cryptocarya sp.
PWOc *ka(m,mʷ)apaR Cryptocarya sp.
PT Muyuw (ka)kamʷeya Cryptocarya sp.
MM Patpatar kamahar Cryptocarya kamahar

5.4. Dillenia sp. (Dilleniaceae)

It was noted in §4.3 that Dillenia species range in size from trees of the lower canopy to tall canopy trees. Although accounts of Bismarcks flora make no mention of the presence of a smaller Dillenia species, there is linguistic evidence that POc speakers knew such a species. Only two Oceanic reflexes have been found, both from Southern Vanuatu, both denoting Dillenia biflora, a tree with reddish brown bark that grows to about 15m (Wheatley 1992: 79). POc *drokol (and PMP *de(k,g)el) is reconstructable on the basis of the Southern Vanuatu data and three non-Oceanic cognates, Blit Manobo (klambug) dakaDillenia megalantha19 (Madulid 2001b: 100), Sundanese səgəl and Javanese ḍəgəl, both ‘Dillenia excelsa’ (Heyne 1950: 1071-1072). Dillenia excelsa is also 10–15 min height.

PMP *de(k,g)el small Dillenia species
POc *drokol small Dillenia species
PSV *ne-dɣol Dillenia biflora’ (Lynch 2004a)
SV Sye ne-tɣul small Dillenia species
SV Anejom̃ ne-cɣel small Dillenia species

5.5. Diospyros spp., ebony, blackwood, B blakwud (Ebenaceae)

Diospyros species are sub-canopy trees around 10 m tall with very hard black wood. Diospyros peekelii is used on New Britain for digging sticks, on New Ireland to make clubs for shark-catching (Floyd 1954, Peekel 1984: 432–433). In the western Solomons the wood of some species is used for carving (Hviding 2005: 142). The fruit of Diospyros species is the persimmon. The fruit of some species are edible and are eaten when other food is scarce (Record 1945).

Although Diospyros species are well known sub-canopy trees, widely mentioned in the literature, the only reconstruction that has proven possible is PSOc *numo. Otherwise only very local cognate sets are found, raising the possibility that the more important species have been introduced since POc times.

PSOc *numo Diospyros spp.’ (Lynch 2004a)
NCV Mwotlap (ɣ[ʋ])nʋm Diospyros ferrea
Proto South Melanesian *n(e,i)mo Diospyros spp.’ (Lynch 2004a) 20
SV Sye nimu(ŋlei) Diospyros ferrea
NCal Jawe (ce)nemo Diospyros austro-caledonicus
NCal Fwâi (ce)nuum Diospyros parviflora
NCal Nemi (ce)nuum Diospyros parviflora
NCal Jawe (ce)nuum Diospyros parviflora

5.6. Euodia spp. (Rutaceae)

Borrell (1989: 130–131) finds 14 species of Euodia21 on Kairiru, ranging from small shrubs a metre high to the medium-sized tree, Euodia elleryana (syn. Euodia tetragona, Melicope elleryana), 15-20 m tall, with a soft white wood that has an unpleasantly musty smell and masses of rose-pink flowers (Peekel 1984: 270).

Peekel reports that larger trunks were used on New Ireland for small outrigger canoe hulls. The wood splits easily and is good for planks (Hviding 2005: 104).

Although POc *bala below is supported by only two reflexes, the notes provided by Kwa’ioloa & Burt (2001: 158) are informative. Kwara’ae bala is a generic term for species which include bala ni kwaru [bala of rock] ‘Euodia elleryana’, bala kwau (lit. ‘white Euodia’ ) or bala fufuri (lit. ‘patchy Euodia’), and simply bala. The authors provide a scientific name only for the first of the three. Apparently excluded from the bala taxon is Euodia hortensis, Kwara’ae foʔoka or foʔaka, which has complex ritual uses both here and at other localities. It is possible that POc *bala also denoted a taxon of Euodia species, the more so as Motu ebala denotes Euodia hortensis, the one species that Kwara’ae bala excludes.

POc *bala taxon including various Euodia spp.’ (?)
PT Motu (e)bala Euodia hortensis’ (Lane-Poole 1925)
SES Kwara’ae bala Euodia spp.

A reasonable (but not the only) interpretation of the data below is that POc *bosi denoted a forest tree with white wood, perhaps Euodia elleryana, as the Meso-Melanesian data suggest, and was reapplied to another tree with white wood, Alphitonia zizyphoides, in southern Vanuatu.

POc *bosi a forest tree with white wood, probably Euodia elleryana
Proto Northwest Solomonic *bosi Euodia spp.
MM Nehan bouh Euodia hortensis
MM Nduke bosi Euodia elleryana
MM Marovo bosi trees of lowland or secondary forest, Euodia spp.
MM Roviana bosi Euodia elleryana’ (Record 1945)
PSV *na-bʷus(Vn) whitewood, Alphitonia zizyphoides’ (Lynch 2001c)
SV Sye na-mpo Alphitonia zizyphoides
SV Kwamera na-pa Alphitonia zizyphoides
SV Kwamera na-pʷesən Alphitonia zizyphoides
SV Anejom̃ na-pʷoθ Alphitonia zizyphoides

Despite the fact that the shrub Euodia hortensis (island musk) is often cultivated and occurs in varieties with acrid-smelling leaves of various shapes, green or yellow, which are used to keep flies away and employed for various medicinal and ritual purposes at many localities (Ivens 1927:362, Peekel 1984: 267-268, Whistler 1991b: 133, Whistler 2000: 119, Wheatley 1992: 91, Hviding 2005: 106; ch.9, §2.1), the only term reconstructed for Euodia hortensis is PCP *usi.

PCP *usi Euodia hortensis
Fij Bauan uði Euodia hortensis
Pn Tongan uhi Euodia hortensis
Pn Samoan usi Euodia hortensis’ (W. McClatchey, pers. comm.)

5.7. Kleinhovia hospita, puzzle tree, guest tree, B namatal (Sterculiaceae)

Kleinhovia hospita is a small to medium-sized tree. In New Ireland it usually grows to 6-10 m tall, but is sometimes as tall as 20-30 m in primary forest (Peekel 1984: 373). A striking feature is its short, branching bole: ‘it sprouts into many trees’ in the words of Kwa’ioloa & Burt (2001: 135).

There is remarkable agreement across sources about its uses, which do not vary much from the Bismarck Archipelago to Vanuatu. Its bast (fibrous inner bark) serves as temporary binding material. The soft light timber provided by young straight branches is used for rafters and internal construction. It also provides excellent firewood, and trees are sometimes ringbarked to turn them into convenient fuel sources. It is reputed to be one of the best woods for starting fires by traditional friction methods (another is the small forest tree Callicarpa pentandra according to Kwara’ae lore) (Powell 1976, Henderson & Hancock 1988: 158-159, Arentz et al. 1989: 94, Wheatley 1992: 225, Kwa’ioloa & Burt 2001: 135, Hviding 2005: 115-116).

Fewer sources mention medicinal uses: the Bola of New Britain use an infusion of the bark, and the leaves are used medicinally in Vanuatu (Powell 1976, Wheatley 1992: 225).

Figure 7.9: Kleinhovia hospita: A, tree; B, shoot with a mature and an old inflorescence; C, leaf.

Most of the terms below are from NCV languages, and they present a minor puzzle. Lynch (2004a) divides them into two groups and makes two resemblant but seemingly irreconcilable PNCV reconstructions, *ma(t,d)aka (Banks, Malakula, Epi) and *matala (Banks, Santo, Malakula, Paama, Efate). There are also MM and SES reflexes, however, and here the terms reflect POc *ma(i)tagaR(a). Lynch’s PNCV *ma(t,d)aka, modified here to *ma(t,d)aga, reflects this unproblematically. This leaves Lynch’s PNCV *matala, which I take to reflect an early borrowing from a language which had lost *-k- but retained *-R- as -l- (the SES languages of Malaita reflect this pattern of reflexes, but there is no good reason to suppose that a language descended from the borrowing source still exists).

POc *ma(i)tagaR(a) Kleinhovia hospita
MM Bola maitaga Kleinhovia hospita’ (Arentz et al. 1989: 94)
MM East Kara mətəkək Kleinhovia hospita
MM Patpatar matakara Kleinhovia hospita
SES Lengo mataga Kleinhovia hospita’ (Henderson and Hancock 1988)
SES Santa Ana magaka Kleinhovia hospita’ (metathesis of PSES *mataga)
PNCV *ma(t,d)aga Kleinhovia hospita
NCV Mwotlap na-mʷtak Kleinhovia hospita
NCV Mota mʷataka tree sp.
NCV Uripiv mʷirek Kleinhovia hospita
NCV Naman midag Kleinhovia hospita
NCV Neve’ei na-mdaŋ Kleinhovia hospita
NCV Tape medek Kleinhovia hospita
NCV Avava midaŋ Kleinhovia hospita
NCV Larëvat medrak Kleinhovia hospita
NCV Nese no-murak Kleinhovia hospita
NCV Lewo (puru)mante Kleinhovia hospita
NCV Baki (buru)minda Kleinhovia hospita

PNCV *matala Kleinhovia hospita’ (Lynch 2004a)
NCV Vurës matal Kleinhovia hospita
NCV Kiai matala Kleinhovia hospita
NCV Nokuku metal Kleinhovia hospita
NCV Sakao ne-ntal Kleinhovia hospita
NCV Araki (vi)mʷarala Kleinhovia hospita
NCV Tamambo (vu)matala tree sp .
NCV Raga matala Kleinhovia hospita
NCV Raga matala Kleinhovia hospita
NCV Port Sandwich madre Kleinhovia hospita’ (J. Lynch, pers. comm.)
NCV Aulua medel Kleinhovia hospita’ (J. Lynch, pers. comm.)
NCV Paamese merai tree sp. used to make bow
NCV Nguna na-matal Kleinhovia hospita
NCV South Efate na-matal Kleinhovia hospita

PMM *p(i,u)lakis and POc *paqu, both ‘Kleinhovia hospita’, are also reconstructable. The forms under ‘cf also’ appear to reflect a variant *paqi which may represent a conflation of the two etyma.

PMM *p(i,u)lakis Kleinhovia hospita
MM Bola bulai Kleinhovia hospita’ (for †pulai)
MM Nehan hule Kleinhovia hospita
MM Babatana vilaki Kleinhovia hospita’ (McClatchey et al. 2005)
MM Nduke valakihi Kleinhovia hospita
SES Bugotu vare Kleinhovia hospita’ (for †vale; perhaps borrowed from a MM language)
POc *paqu Kleinhovia hospita
MM Tolai vau Kleinhovia hospita’ (Record 1945)
MM Roviana paɣo Kleinhovia hospita’ (Record 1945)
NCV Lakon vʋɣ-vʋɣ Kleinhovia hospita’ (François 2004)
Fij Wayan vau Kleinhovia hospita
cf. also:
MM Mono-Alu (la)hai Kleinhovia hospita
SES Lau fai-fai Kleinhovia hospita
SES Kwara’ae fae-fae Kleinhovia hospita
SES Kwaio fae-fae tree sp.

5.8. Litsea spp. (Lauraceae)

Numerous Litsea species of varying statures occur in the Bismarcks and the Solomons. If POc *lowaŋa, reconstructed below, did indeed denote a particular species of Litsea, then O’Collins & Lamothe (1989) indicate that it was one large enough to make a canoe hull. Of the species listed by Peekel (1984: 191), two qualify: Litsea kauloensis and Litsea dielsiana, each 15-20 m tall.

The reconstruction of PCEMP/POc *lowaŋaLitsea sp.’ is made possible by the CMP reflexes recorded by Verheijen (1990: 224): Kepo, Rembong (both CMP) lowaŋLitsea sp.’. Although Nakanai loagaGmelina sp.’ is glossed as another species, both are used for canoe hulls, and the Nakanai term is almost certainly cognate with Nyindrou lowaŋ. Nakanai also provides us with the final vowel for the reconstruction; that there was a final vowel can be inferred from the fact that final consonants are lost in Nyindrou and in the CMP languages from which reflexes are drawn.

PCEMP *lowaŋa Litsea sp.
POc *lowaŋa Litsea sp.
Adm Nyindrou lowaŋ Litsea sp.’ (O’Collins and Lamothe 1989)
MM Nakanai loaga Gmelina sp.

5.9. Myristica spp., wild nutmeg, B nandae (Myristicaceae)

The most famous Myristica species is the nutmeg, but its easternmost extent is West Papua (Warburg 1899: 57). Among the species of wild nutmeg which grow in the Bismarcks are Myristica schleinitzii, a small foreshore tree 3-6 m tall, and Myristica fatua, a sub-canopy tree reaching 20m on New Ireland (Peekel 1984: 185). Myristica fatua is a canopy tree in some areas, e.g. in Kwara’ae country. Both species prefer a damp environment and have stilt roots even in dry locations (Paijmans 1976: 37). POc *(dr,d)aRa(q,k)a presumably denoted at least one of these two species and perhaps a taxon including a number of Myristica species.

The soft wood of both species makes planking for indoor use and good firewood in locations from New Britain to Vanuatu (Floyd 1954, Powell 1976, Wheatley 1992:172-174, Kwa’ioloa & Burt 2001: 120, 167).

With regard to reconstructing a PROc term for Myristica fatua Lynch (2004a) writes:

The Banks and Raga forms suggest *draRaka, other NCV languages something like *draRaqi, while the SV forms point rather to PSOc *d(r)ani, *d(r)aRani or *d(r)aqani. Ross Clark (pers. comm.) has suggested a possible connection to words meaning ‘blood’, from POc *draRaq: Wheatley (1992: 172) refers to the ‘dark red exudate’ of the inner bark, and Codrington relates Mota naraa to nara ‘blood’. The S Efate form is also homophonous with ‘blood’, and the SV forms almost so (PSV *n-da(q,V), *n-da(a)-). The form may thus be based on the POc term for ‘blood’ [*draRaq], and be something like *draRaq-(n)(i,a).

To Lynch’s Southern Oceanic cognate set we add Western Oceanic and Kwara’ae (SES) items. Reconstructing a POc etymon from these data is tricky, as they do not lead to an unambiguous reconstruction, yet they probably all reflect a single etymon.

Lynch’ s discussion points to three phonological questions:

  • the third consonant: was it POc *k or *q?
  • the third vowel: was it POc *a or *i?
  • what is the source of PSV *n?

To these we can add a fourth not raised by Lynch’s data:

  • the initial consonant: was it POc *dr or *d?

I address the last question first. All reflexes except Muyuw, Lihir and Kwara’ae point to either *dr- or *d-. Muyuw a-yayak and Lihir lala, however, reflect initial POc *r- or *R-. These probably reflect assimilation to the medial reflex of *-R-, so they do not help us to disambiguate the initial. Distinguishing reflexes of *d from those of *dr is difficult, because *d was a very low-frequency POc consonant: for some languages we have no reflex of *d, and in some others it merges with *dr. The one language that helps us with this disambiguation is Kwara’ae, where ka-kalaʔa at first sight seems to reflect POc *g-. However, a small number of SE Solomonic lexical items reflect POc *d (but apparently not *dr) as if it were *g, and this is evidently one of them. This suggests that the initial consonant was *d- and that resemblances to the word for ‘blood’ are accidental, but perhaps amplified by folk-etymologising, since the blood-like colour of Myristica sap is widely recognised.22

The second question to be addressed concerns the third consonant: was it POc *k or *q? As Lynch notes, the Banks and Raga reflexes (the first six NCV reflexes below) point to *k. So does Muyuw a-yayak. Other reflexes point to *q. If changes in form took place as a result of folk-etymologising, then reflexes of *q may be due to reflexes of POc *draRaq ‘blood’, leaving *k as the more likely proto-consonant.

The other two questions-was the third vowel *a or *i? what is the source of PSV *n? - may be taken together. Four items below, Kwara’ae ka-kalaʔa, Vera’a daraɣa, Mota na-raɣa and Raga a-oaɣa suggest that the vowel was *a, i.e. they point to POc *(dr,d)aRaka. On the other hand NE Ambae dadai and Uripiv drari reflect a final *-i in *(dr,d)aRaq(a)-i, and PSV *na-(dr,d )ani reflects a final *-ni in *(dr,d )aRaq(a)-ni (these are languages in which both *R and *q are lost). The altemation between final *-i and *-ni is suggestive, as both are reflected in variants of the so-called associative: (non-specific possessor) construction. Thus ‘blood of tree’ would have been expressed in POc as *(dr,d)aRaq i kayu or *(dr,d)aRaq ni kayu, depending on whether *(dr,d)aRaq was directly or indirectly possessed; either *i or *ni has been generalised to both in various daughter-languages (Ross 1998b). The suggestion is that *-i- and *-ni_ are accretions brought about by the truncation of something like ‘blood of tree’, itself an outcome of folk etymology, and that they are therefore not part of the POc reconstruction.

The reconstruction that emerges from the discussion above is POc *daRaka, but there is enough speculation above to commend prudence, and so I offer POc *(dr,d)aRa(q,k)a below.

POc *(dr,d)aRa(q,k)a wild nutmeg, Myristica sp.
PT Muyuw (a)yayak Myristica schleinitzii
MM East Kara de Myristica fatua
MM Lihir lala Myristica sp.’ (Burley 2006)
SES Kwara’ae ka-kalaʔa Myristica fatua

PNCV *(dr,d)aRa(q,k)(a,i) wild nutmeg, Myristica sp.
NCV Mota na-raɣa nutmeg
NCV Mwotlap na-d[a]yaɣ Myristica fatua’ (François 2004b)
NCV Vera’a daraɣa Myristica fatua’ (François 2004b)
NCV Vurës daraɣ Myristica fatua’ (François 2004b)
NCV Mwesen (wo)na-raɣ Myristica fatua’ (François 2004b)
NCV Raga (a)oaɣa Myristica fatua
NCV Ambae dadai Myristica fatua
NCV Uripiv drari Myristica fatua
NCV South Efate n-ra Myristica fatua
PSV *na-(dr,d)ani wild nutmeg, Myristica fatua’ (Lynch 2004a)
SV Sye na-nre Myristica fatua
SV Lenakel ne-tan Myristica fatua
SV Kwamera n-tan Myristica fatua
SV Anejom̃ na-jeñ Myristica fatua

5.10. Parinari spp., putty nut (Chrysobalanaoeae)

Finding one’s way through the maze of synonymous scientific names for Bismarcks species of the genus Parinari is difficult, but it appears that in NW Island Melanesia there are two similar species which were perhaps treated as a single POc taxon, *(q,k)atita:23

  • Parinari laurina (syn. Cyclandrophora laurina, Atuna racemosa) and
  • Parinari glaberrima (syn. Maranthes corymbosa, Parinari corymbosa, Parinari griffithiana) (Figure 7.10, left)24

Parinari laurina seems to be more common in the Bismarcks, Parinari glaberrima in the Solomons. Both are small to medium-sized sub-canopy trees, 10–20 m tall. Their large nuts have a hard shell. After it has been broken, the tough kernel of the fruit is rubbed over a rough surface to produce small crumbs of putty-like mash, and the resulting sticky putty is used to caulk plank canoes, to seal and repair cracks in dugouts, and to fasten shell inlays to wood carvings. After drying, this coating forms a firm, watertight and breakable-resistant layer (Holdsworth & Wamoi 1981, Peekel 1984:202-203, Henderson & Hancock 1988:230-232, Ohnemus 1998, Kwa’ioloa & Burt 2001: 165, Hviding 2005: 147).

In the Carolines a decoction of the pericarp of the Parinari laurina fruit is used for painting canoes red (Christian 1899: 328). In Fiji its long straight branches are used for canoe paddles and as houses rafters. Its leaves are used to fill in the outer walls of houses (Capell 1941). In the Solomons the bark provides a medicine taken against diarrhoea or dysentery (Henderson & Hancock 1988: 232).

Reconstructing the POc term for the putty nut is a little tricky, as Chowning (2001: 76) mentions in a footnote, because Oceanic languages appear to reflect three related forms: *qatita, *katita and *tita. The Admiralties forms below may reflect either POc *katita or

Figure 7.10: Left Parinari glaberrima: A, tree; B, shoot bearing one mature and one old inflores- cence; C and D, nut. Right Caryota rumphiana: A, tree; B, leaflet; C, portion of inflorescence; D, fruiting inflorescence; E, fruit; F, larvae of Rhynocaphorus beetle in a rotting trunk.

POc *qatita. Of the Western Oceanic forms, Wogeo kətita and Kara katita reflect *katita, whilst the rest may reflect either *katita or *qatita. On the simplest interpretation of the data I reconstruct PWOc *katita. The Eastern Oceanic forms, on the other hand, reflect *qatita. POc *(q,k)atita below reflects this ambiguity.

POc *(q,k)atita the putty nut, probably Parinari laurina and Parinari glaberrima’ (ACD: *qatita)
Adm Likum ketik putty nut, Parinari laurina
Adm Drehet ketik putty nut, Parinari laurina
Adm Lou kerit putty nut, Parinari laurina
PWOc *katita the putty nut, probably Parinari laurina and Parinari glaberrima
NNG Kove atita putty nut, Parinari laurina
NNG Wogeo kətita putty nut, Parinari laurina
NNG Kilenge atita putty nut, Parinari laurina
MM East Kara katita putty nut, Parinari laurina
MM Tolai katita putty nut, Parinari laurina
MM Tinputz acic putty nut, Parinari laurina
MM Teop asita putty nut
PEOc *qatita the putty nut, probably Parinari laurina and Parinari glaberrima
SES Baegu saia putty nut
SES Kwara’ae saia putty nut, Parinari glaberrima
SES Sa’a saie putty nut
SES Lau saia putty nut, Parinari glaberrima
SES Kwaio laia putty nut
SES ’Are’are raia putty nut
Mic Carolinian ais putty nut, Parinari laurina
Mic Chuukese ayis Parinari tree
Mic Woleaian yaise tree sp. with fragrant fruit
Mic Ponapean ays Parinari tree

Blust (ACD) reconstructs doublets *qatita and *qarita for this item. Supporting data for medial *-r-, however, are drawn entirely from Admiralties languages and Mussau, a distribution which does not justify a POc reconstruction on the criteria set out in ch. 1, §3.2.3. Instead, it seems likely that the forms listed below which appear to reflect *qarita are the result of borrowing(s) from a language or languages that reflects (or reflected) POc *-t- as -r-. There are a number of these in the Admiral1cies: Lou, Penchal, Baluan, Lenkau, Pak, Koro, Nali, Lele and Ponam (Ross 1988: 322) (among the data above only Lou has -r- in a directly inherited reflex of *qatita).

Among directly inherited Admiralties reflexes of *qatita, those in Likum, Drehet and Lou have initial k-. The reflexes below, however, agree not only in appearing to reflect *-r- but also in loss of the initial consonant. This suggests that they may all reflect a single early borrowing of a form *arita.

Adm Mussau arita putty nut
Adm Nyindrou alik putty nut, Parinari laurina
Adm Nauna alit putty nut, Parinari laurina
Adm Penchal alit putty nut, Parinari laurina
Adm Pak ehir putty nut, Parinari laurina
Adm Nali n-alit putty nut, Parinari laurina
Adm Ere arit putty nut, Parinari laurina
Adm Titan alit putty nut, Parinari laurina
Adm Sori-Harengan ahiʔ putty nut, Parinari laurina
Adm Leipon yerit putty nut, Parinari laurina
Adm Loniu eit, aʔat putty nut, Parinari laurina

A number of Meso-Melanesian reflexes lack the initial syllable, reflecting a possible PMM altemant *tita.25 For clarity’s sake these are listed separately below. There are also two SE Solomonic reflexes of *tita, in Gela and Bugotu, but both languages are prone to borrow from NW Solomonic (and thus Meso-Melanesian) neighbours.

Attempting to account for *tita Blust (ACD) suggests that the initial syllable has been irregularly lost. He puts forward two possible reasons for such a loss. First, POc roots were predominantly disyllabic, and this is true of many daughter-languages: this might favour foreshortening. Second, the POc common article was *a or *na. If initial *k- or *q- was lost from a reflex, then the resulting initial *a- could be reanalysed as part of the article. A third possible reason is offered here, namely that *ka- was reanalysed as the ‘tree’ prefix *ka-, leaving *tita as the name of the tree (ch.2, §7.1.2).

PMM *tita the putty nut, probably Parinari laurina and Parinari glaberrima’ (Chowning 1963)
MM Nakanai tita Parinari glaberrima
MM Tolai tita Parinari laurina
MM Nehan tita Parinari laurina
MM Petats tic Parinari glaberrima
MM Teop tita Parinari glaberrima
MM Varisi sita putty nut, Parinari glaberrima’ (W. McClatchey, pers. comm.)
MM Simbo tita Parinari laurina; gum, glue
MM Nduke tita Parinari glaberrima
MM Marovo tita Parinari glaberrima
MM Roviana tita Parinari glaberrima’ (Henderson and Hancock 1988)
SES Bugotu tita putty nut
SES Gela tita Parinari sp.
cf. also:
MM Babatana lita Parinari glaberrima

The cognate set below appears at first sight to be related to Baegu, Kwara’ae, Sa’a and Lau saia above, but saia is the regular reflex of POc *qatita, since *q- and *-t- are both deleted, and s- is a regular accretion before the resulting initial a-, corresponding with Kwaio l- in laia and ’Are’are r- in raia (Frantisek Lichtenberk 1988). These changes are quite different from those regularly reflected in PCP.

PCP *sea tree, Parinari insularum’ (Milke (1961): POc)
Fij Rotuman sea tree sp.
Fij Bauan sea Parinari insularum
Pn Tongan hea Parinari insularum
Pn East Futunan sea Parinari insularum
Pn East Uvean hea Parinari insularum
Pn Samoan sea Parinari insularum

It is not clear what connection (if any) the form below has with those above.

POc *maRakita the putty nut, probably Parinari laurina and Parinari glaberrima
MM Mono-Alu malakita putty nut, Parinari glaberrima’ (W. McClatchey, pers. comm.)
Fij Wayan mākita forest tree with large seeds used to caulk canoes, Parinari laurina
Fij Bauan makita putty nut, Parinari laurina

5.11. Palms

5.11.1. Caryota rumphiana, black palm, fishtail palm, TP waillimbum (Arecaceae)

Caryota rumphiana is the only palm in NW Island Melanesia with bipinnate fronds, i.e. the leaflets on either side of the midrib themselves have a central rib and are leaf-like (Figure 7.10, right). Caryota rumphiana stands 10–20 m high and has fruit the size of a cherry which hang in bunches from the top of its blackish trunk (Peekel (1984), Kwa’ioloa & Burt 2001: 187).

Throughout NW Island Melanesia, the trunks are split to make floorboards (Floyd 1954, Henderson & Hancock 1988: 150, Arentz et al. 1989: 93, Scales n.d. McEldowney 1995, Margetts 2005a). The wood is also used on New Britain for axe handles, clubs, bows and spears (Floyd 1954, Powell 1976). The pith from the young trunk is sometimes eaten on New Britain, and fed to pigs in the Solomons.

Henderson & Hancock (1988: 150) describe how inland dwellers in the Solomons use a felled Caryota rumphiana trunk to farm larvae of a large beetle of the genus Rhynocaphorus by cutting notches at 2 m intervals along it, harvesting the larvae and pupae 3–4 months later. For some households the larvae are a major source of protein, although others find the accompanying taste of the rotting palm core rather offensive.

The Oceanic data below were assembled by Blust (ACD). The reconstruction of PCEMP *bual(a)Caryota sp.’ is based on these data and on W Sumba ʔwuola, E Sumba (both CMP) ʔwualaCaryota mitis’ (Verheijen 1990: 199).

PCEMP *bual(a) Caryota sp.
POc *[bual]bual species of palm used for making spears and bows; palm-wood spear or bow, probably Caryota sp.’ (ACD)
Adm Lou (si)pua black palm
PT Tawala bua-bua smalll tree fern, used for spears
PT Saliba bua-bua k tree, used for spears and sticks
MM Lihir buer fern sp.
MM Roviana buala large kind of bow
MM Simbo buala war bow
SES Sa’a pue-pue a palm used for making bows, combs, heavy spears’; ‘heavy palm-wood spear
SES Sa’a pue-pue a palm used for making bows, combs, heavy spears’; ‘heavy palm-wood spear
SES Ulawa pua-pua a palm used for making bows, combs, heavy spears
Pn Hawaiian pua arrow, dart, sometimes made from flower stalks of sugarcane

PMP *katipa(l,n) below is reconstructed on the basis of the Oceanic data below and of Hanunuo, Mangyan (WMP) katipanCaryota cumingii’ (Madulid 2001a: 364) and Wandamen (EMP) kasira ‘black palm sp.’ (Smits & Voorhoeve 1992: 222).

PMP *katipa(l,n) a palm with black wood, Caryota sp.
POc *kati(p)al a palm with black wood, Caryota sp.
PT Tawala kahiala Caryota sp.
PT Dobu kasiala Caryota sp.
MM Tinputz kacan Caryota sp.
MM Marovo kacuele palm, Drymophloeus sp., black wood used for bows and spear tips

Tiny though the cognate set below is, the membership of its two members in different primary subgroups of Oceanic justifies a POc reconstruction.

POc *j(o,u)abo Caryota sp.
Adm Baluan soap Caryota rumphiana
MM Bali tuabo Caryota sp.’ (Hide 1985)

5.11.2. Licuala spp., fan palm (Arecaceae)

The leaves of the small palm Licuala ramsayi (syn. L. muelleri, L. peekelii), growing to about 5-10 m, serve for roofing in parts of the Bismarcks (Powell 1976, Peekel 1984: 58). The Nakanai use them to wrap megapode eggs (A. Chowning, pers. comm.). The palm also serves decorative purposes: in the Ninigo Islands it is planted as an ornamental shrub (Sorensen 1950), and various writers note that its leaves are used for personal decoration.

POc *piRu denoted one or more Licuala species, a fan palm. As French-Wright (1983: 208-209) and Chowning (2001: 84) note, in Fijian and the Polynesian languages its reflexes denote the fan palm Pritchardia pacifica, found only in Fiji and Polynesia.26 This is an instance of an established name being given to a new-found species as Oceanic speakers moved eastward.

On the basis of the non-Oceanic data supporting Blust’s reconstruction of PMP *biRuʔ (ACD), we would expect the POc form *piRu, and this is reflected everywhere except in northern Vanuatu, where a local form *piloqi is reflected. NCV forms sporadically retain a POc final consonant with an added *-i, regularly lost in Volow, Mota and Merlav, and so the final of these items may reflect POc *-q, a possibility recognised in the reconstruction of POc *piRu(q).

PMP *biRuʔ fan palm, Licuala rumphii’ (ACD)
POc *piRu(q) fan palm, Licuala sp.’ (ACD; Chowning 2001)
NNG Kove pilu Licuala sp.
MM Nakanai vilu-vilu Licuala sp.
MM East Kara fi Licuala lauterbachii
MM Halia hil Licuala sp.
MM Solos hin Licuala sp.
MM Mono-Alu hiuru Licuala lauterbachii’ (W. McClatchey, pers. comm.)
PEOc *piRu(q) fan palm, umbrella palm’ (Geraghty 1990)
SES Gela vilu species of palm with umbrella-like leaves
SES Lau filu umbrella palm
SES Kwaio filu wild palm species used to make bows
SES Kwara’ae filu Pritchardia pacifica
SES Kwara’ae filu tali Licuala lauterbachii
SES Sa’a hilu a ridge covering of sago-palm leaves, laid on flat
SES ’Are’are hiru a palm tree with digital leaves, used for making bows
SES Arosi hiru a palm sp. used in making a war-bow
NCV Volow (n-ye)ploɣ Licuala sp., Pritchardia pacifica’ (François 2004b)
NCV Mota viloɣ an umbrella palm; a frond of that palm used as an umbrella
NCV Dorig (dã)vlʋ Licuala sp., Pritchardia pacifica’ (François 2004b)
NCV Merlav (dʋ)vlʋɣ Licuala sp., Pritchardia pacifica’ (François 2004b)
SV Sye (lu)vor Pritchardia pacifica
NCal Xârâcùù pii a kind of palm tree
PCP *viu fan palm, umbrella palm, Pritchardia pacifica
Fij Bauan viu Pritchardia pacifica
Pn Tongan piu Pritchardia pacifica
Pn Niuean piu Pritchardia pacifica
Pn East Uvean piu Pritchardia pacifica
Pn East Futunan piu fan palm
Pn Samoan piu Pritchardia pacifica
Pn Tokelauan piu Pritchardia pacifica

6. The shrub layer

The shrub layer is usually patchy because of lack of sunlight. A few small trees, up to a height of around 15m, grow here, as well as Calamus (§6.3.1), bamboos (ch.13, §3.1) and gingers (Henderson & Hancock 1988: 320).

6.1. Woody shrubs

6.1.1. Abroma augusta (Sterculiaceae)

Abroma augusta is a small shrub 1-2m tall, more common in the highlands of New Guinea than in lowland forests (French 1986, Peekel 1984: 373). On Malaita its white bark is used to make hanging baskets (Kwa’ioloa & Burt2001: 160). In parts of New Britain its bast provides rope for pig nets and for lashing house components and material used in making clothing, and bags (Powell 1976, Lentfer 2003).

Reflexes of POc *wasi-wasiAbroma augusta’ are found with this meaning only in Near Oceania. In Remote Oceania the name denotes a large forest tree, Sterculia vitiensis (ch. 11, §2.6), which does not occur in NW Island Melanesia. Will McClatchey (pers. comm.) suggests that the change in denotation was mediated by the fact that Sterculia vitiensis is also a source of fibre.

POc *wasi-wasi Abroma augusta
MM East Kara (ka)us-vas Abroma augusta
MM Patpatar was-was Abroma augusta
MM Nehan ase-is Abroma augusta
SES Kwara’ae kʷasi-kʷasi Abroma augusta
PROc *wasi-wasi Sterculia vitiensis’ (from data in Wheatley 1992, Lynch 2004a)
NCV Ambae wah-wah Sterculia vitiensis
NCV Tangoa (vitu)vaha Sterculia vitiensis
NCV Raga wahi-wahi Sterculia vitiensis
NCV Apma wah-wah Sterculia vitiensis
NCV Tape (vən)woso-wos whitewood
SV Sye wo-wo Sterculia vitiensis
SV Lenakel nə-vha-vha Sterculia vitiensis27
SV Anejom̃ n-woθ-waθ Sterculia vitiensis
Fij Buca Bay waði-waði Sterculia vitiensis’ (J. Parham 1972; Buca Bay is in Vanua Levu)

6.1.2. Angiopteris evecta (syn. Angiopteris erecta), mule’s foot fern (Marattiaceae)

The plant with the largest fronds (1.5m long) on New Ireland, Angiopteris evecta is a large fern common in shady inland forest. The fronds rise from a massive rootstock, fleshy and moist, but fragrant when dry (Peekel 1984: 30). Because of their moisture, the Kwara’ae lay them out around garden boundaries to soften the soil in the belief that this will improve the quality of the taro (Kwa’ioloa & Burt 2001: 211).

PCP *nas(e,i) edible roots of certain plants ?’ (POLLEX2)
Fij Bauan naði the greater roots of the yaqona plant
PPn *nas(e,i) giant fern, Angiopteris evecta, with edible root
Pn Samoan nase the giant ferns Angiopteris evecta and Marattia fraxinea
Pn Marquesan nahe, nahi root eaten in times of scarcity
Pn Rarotongan naʔe Angiopteris evecta
Pn Tahitian nahe Angiopteris evecta

6.1.3. Donax cannaeformis (syn. Clinogyne grandis, Maranta grandis, Thalia cannaeformis, Actoplanes canniformis, Donax arundastrum) (Marantaceae)

A leafy shrub, Donax cannaeformis has a rhizome from which rise erect smooth stems 1.5-3 m tall with spreading branches about 85cm long. It has elliptical leaves often with yellowish-white patches and white flowers and fruits, and grows in damp locations (Peekel 1984: 111). Its range is from SE Asia to the Solomons.

As the glosses below indicate, its stems are used in thatching, basket-making and as armlets.

PMP *niniq plant sp., Donax cannaeformis, used as material for making baskets’ (Blust 1989; ACD)
POc *nini(q) shrub, Donax cannaeformis
MM Bola (natala)nini Donax cannaeformis
MM Marovo nina Donax cannaeformis
MM Ririo nina Donax cannaeformis’ (W. McClatchey, pers. comm.)
MM Babatana nine Donax cannaeformis’ (W. McClatchey, pers. comm.)
SES Gela nini sp. of bush used in wristlets
SES Lau nini sp. of shrub; stems used in thatching

6.1.4. Garcinia spp. (Clusiaceae)

Garcinia species grow in all sizes from small shrubs to tall canopy trees. The best known species is G. mangostana, the mangosteen, which grows up to 25 m high, but this is not indigenous to Oceania. The only indigenous species reported by Peekel (1984: 376-377) in the Bismarcks is Garcinia novo-guineensis (syn. Garcinia warburgiana), a tree just 5-10 m tall with small white flowers. It contains, especially in the roots, yellow latex which is used on New Ireland for painting ancestor pictures. This yellow is greenish and lighter than that from Curcuma (ch.13, §5.1).

Another species found in NW Island Melanesia is Garcinia pseudoguttifera (syn. Garcinia pancheri), a tree up to 25m high (Wheatley 1992: 112). It is not found in the Bismarcks but occurs from Bougainville to Tonga. It has edible fruit and displays the considerable variation that reflects former cultivation (Walter & Sam 2002). Taller Garcinia species appear to play a greater role in the rain forests of Bougainville than of the Bismarcks (Mueller-Dombois & Fosberg 1998: 61).

Wood from an unnamed Garcinia species is used for axe handles on New Britain and for rafters both there and in the Admiralties (Floyd 1954, Arentz et al. 1989, O’Collins & Lamothe 1989).

Very little information is available to help us determine the denotata of the species below, but it is a reasonable inference that POc *bulu denoted the small Garcinia novo-guineensis, as this is present in the Bismarcks (and is the gloss of the Motu reflex).

POc *bulu Garcinia sp., perhaps Garcinia novo-guineensis
PT Motu bio-bio Garcinia novo-guineensis’ (Lane-Poole 1925) 28
SES Kwara’ae (ʔai) bulu Diospyros maritima
SES Lau (ʔai) bulu Diospyros maritima
NCV Vera’a wu-wul Garcinia pancheri’ (François 2004b)
NCV Lakon vu-vul Garcinia pancheri’ (François 2004b)
Fij Bauan bulu Garcinia spp.’ (Keppel et al. 2005)

The denotatum of PWOc *tabun was probably a large canopy tree. Muyuw tob is a large canopy tree with stilt roots, an apt description of Garcinia latissima (Conn & Damas 2006). Tolai tabu-tabun is glossed Garcinia scaphopetala, which grows to 30 m.29

PWOc *tabun Garcinia sp.
PT Muyuw tob large (25m) Garcinia sp. with aerial roots
MM Tolai tabu-tabun Garcinia scaphopetala’ (Record 1945)

The tree designated by PSV *n-mobʷol was evidently also a large species, as Garcinia sessilis and Garcinia platyphylla are both canopy trees.

PSOc *ma(b,bʷ)ola Garcinia sp.’ (Lynch 2004a)
NCV Mota maploa a tree, with smooth scented leaves and bark
PSV *n-mobʷol Garcinia sp.’ (Lynch 2004a)
SV Sye mompol Garcinia sessilis
SV Anejom̃ n-mopʷol(hat) Garcinia platyphylla

6.1.5. Phaleria spp. (Thymelaeaceae)

Peekel (1984: 393) describes Phaleria coccinea as a climbing shrub, 2-4m high, with white flowers and red berries. No term is reconstructable earlier than PEOc *sinu, glossed ‘shore tree with scented white flowers’ by Geraghty (1983). The Bauan gloss points to a taxon of shrubs whose sap causes irritation, including species of Phaleria, an inference supported by the Tongan and Samoan glosses.

If the NW Solomonic items under ‘cf. also’, all glossed Cominsia gigantea (Marantaceae) (a leafy shrub), are cognate, then POc *jinu is reconstructable but with uncertain denotation.

PEOc *sinu taxon of shrubs whose sap causes irritation, including species of Phaleria’ (Geraghty 1983: 139)
SES Gela sinu k.o. shore tree
Fij Wayan sinu small coastal tree, Excoecaria agallocha, with acrid milky sap, capable of blinding
Fij Bauan sinu generic name for several trees whose sap is irritating, including Phaleria spp.
Fij Rotuman huni flowering bush, Phaleria disperma’ (metathesis: Pn borrowing)
PPn *sinu Phaleria sp.
Pn Tongan huni flowering bush, Phaleria disperma’ (metathesis)
Pn Niuean huni various kinds of plants with clustered flowers’ (metathesis)
Pn East Futunan sinu a flowering shrub, Hoya bicarinata
Pn West Futunan sinu a tree whose sap is said to cause blindness
Pn Ifira-Mele sinu a tree with irritating sap
Pn Emae sinu a tree sp.
Pn Samoan suni Phaleria disperma; taxon of flowering shrubs, inc. Phaleria spp., introduced Ixora spp. and Hoya australis’ (metathesis) (Whistler 2000: 198-199)
cf. also:
MM Babatana zi-zinu (mesara) Cominsia gigantea’(McClatchey et al. 2005)
MM Nduke zinu Cominsia gigantea
MM Roviana zinu Cominsia gigantea
MM Marovo sinu Cominsia gigantea

6.1.6. Semecarpus forstenii, poisonwood, B naolasi, posentri, posenwud (Anacardiaceae)

Semecarpus forstenii is a shrub or small tree 3-10m tall, with a corrosive black sap which destroys skin and inflicts painful wounds. Peekel calls it ‘[t]he most feared tree in the Bismarck Archipelago’ (Peekel 1984: 328, Kwa’ioloa & Burt 2001: 121-122).

Semecarpus vitiensis (syn. S. laxiflora) is a medium-sized tree up to 25 m in height, but it has a black sap with similar effects to that of S.forstenii (Wheatley 1992: 38).

The distribution of the two species seems to be complementary: Semecarpus forstenii in the Bismarcks and the Solomons and the larger Semecarpus vitiensis in Vanuatu and Fiji.30 For this reason I infer that POc *walasi denoted Semecarpus forstenii. POc *lasi is listed by Tryon (1994) as ‘Antiaris toxicaria’, a gloss maintained by Lynch (2002f), but this seems incorrect in view of the reflexes listed here.31

The presence of -i- in Kwaio and Kwara’ae kʷailasi and Lau koilasi probably reflects a folk etymology which interprets the first syllable as kʷai ‘river, water’ (< POc *waiR), but there is no other evidence to suggest that this is the origin of POc initial *wa-. Indeed, it is possible that this folk etymologising accounts for the loss of *wa- in a number of reflexes.

PMP *laji tree sp. with poisonous sap, Antiaris toxicaria (?)’ (Blust 1986; ACD) 32
POc *walasi tree sp. with poisonous sap, Semecarpus forstenii
Adm Loniu walas a long seagrass which grows on sandy area near shore
NNG Takia walas tree sp.
PT Molima wenasi Semecarpus sp.
MM Patpatar (i)walas Semecarpus forstenii
MM Tolai ola Semecarpus forstenii
MM Gao na-ulahi Semecarpus forstenii’ (W. McClatchey, pers. comm.)
MM Maringe n-olahi Semecarpus forstenii’ (W. McClatchey, pers. comm.)
SES Kwaio kʷailasi Semecarpus sp.
SES Lau koilasi Semecarpus sp.
SES Kwara’ae kʷailasi Semecarpus forstenii
SES Arosi warasi sp. of tree with edible yellow fruit
SES Sa’a lasi tree sp. with juice causing sores
PSOc *walasi Semecarpus vitiensis
NCV Mwotlap leh Semecarpus vitiensis
NCV Mota las tree sp.
NCV Ambae walahi Semecarpus vitiensis
NCV Nokuku aulasi Semecarpus vitiensis
NCV Kiai olasi Semecarpus vitiensis
NCV Araki (vi)olas Semecarpus vitiensis
NCV Tamambo (vu)alasi tree sp.
NCV Sakao elai Semecarpus vitiensis
NCV Raga walahi Semecarpus vitiensis
NCV Labo na-walas Semecarpus vitiensis
NCV South Efate n-las Semecarpus vitiensis
PSV *na-ɣilas Semecarpus vitiensis’ (Lynch 2001c)
SV Sye no-ule Semecarpus vitiensis
SV Lenakel ni-lha Semecarpus vitiensis
SV Anejom̃ ne-ɣlaθ Semecarpus vitiensis
NCal Pije wãnit Semecarpus vitiensis
NCal Iaai (i-o)unic Semecarpus vitiensis
NCal Nêlêmwa wââric Semecarpus vitiensis

6.2. Leafy shrubs

The only wild leafy shrub for which a reconstruction has been made is Hornstedtia lycostoma. The genera Alpinia and Heliconia also belong here, but no reconstruction of a name for an Alpinia species is supported by the data. Heliconias are also cultivated, and are treated in ch.13, §6.5.

6.2.1. Hornstedtia lycostoma (syn. Hornstedtia scottiana) (Zingiberaceae)

Hornstedtia lycostoma is a leafy shrub, 3-6m tall, a tall wild ginger with long leaves and red flowers that issue directly from the stem (Figure 7.11, left). The edible seeds are sweet and are gathered especially by children who sometimes eat so many that they become constipated (Powell 1976, Peekel 1984: 105-106, Kwa’ioloa & Burt 2001: 195 Hviding 2005: 110).

POc *dali-dali Hornstedtia lycostoma
MM Patpatar dal-dal Hornstedtia lycostoma
MM Tolai (ta)dal-dal Hornstedtia lycostoma
MM Ramoaaina dal-dal a plant
MM Maringe da-dali Hornstedtia lycostoma’ (Henderson and Hancock 1988)
SES Kwara’ae ka-kali Hornstedtia lycostoma’ (Henderson and Hancock 1988)
Fij Wayan dali-dali Polyscias sp.

6.3. Climbers and epiphytes

6.3.1. Calamus spp., rattan, lawyer cane, TP kanda, P loeaken (Arecaceae)

There is a sense in which rattan could be assigned to the canopy, as it uses canopy trees as hosts and sometimes climbs as high as 50 m. At the same time, it is not a tree and it is convenient to treat it alongside other non-treelike plants.

Peekel (1984: 61) describes two very similar species of rattan, Calamus hollrungii (Figure 7.11, right) and Calamus ralumensis. They are spiny climbing palms from the vines of which curved thorns protrude to attach it to the host. Calamus hollrungii is recorded throughout NW Island Melanesia, whereas Calamus ralumensis is reported only by Peekel and only on the Gazelle Peninsula of New Britain. This raises the possibility that it is identical with Calamus stipitatus, which occurs throughout the Solomons and is described as similar to Calamus hollrungii but having longer, narrower leaflets and a somewhat thinner vine (Henderson & Hancock 1988:208-211, Hviding 2005: 134, 147). Rattan appears not to have been present in Remote Oceania until recently.

Reports from the north coast of New Britain say that pieces of rattan are used for arrowheads, for the binding on arrows, for adzes, for bowstrings and bow bracers, in boats for lashings, bindings and braces and for the anchor cable, in houses for tying and plaiting, and for personal adornment as armbands and armlets, belts, necklaces and headbands (Floyd 1954, Powell 1976). A similar range of uses of the cane is reported elsewhere. Hviding reports that split lengths of Calamus stipitatus are used for sewing roofs and sewing sago-leaf panels in house construction. Unsplit lengths make ropes for heavy tasks like pulling a dugout canoe from its construction site down to the beach. Calamus hollrungii has similar uses, according to Kwa’ioloa & Burt (2001: 205). Hviding mentions that it is used to make tongs to pull items from the hot stone oven. Other parts of the plant are also used. The Bola make wall insulation from the leaves, sometimes eat the young shoots, and use the sap for various medicinal purposes. At Kwara’ae the thorns are used as tattooing needles.

POc *qu(w)e33 presumably denoted all of the two or three species mentioned above.

PAn *quay rattan, Calamus sp.’ (ACD)
POc *qu(w)e rattan, Calamus spp.’ (Grace 1969)
PT Gapapaiwa kuvei rattan
PT Wedau uwe rattan
PT Bwaidoga uwe rattan
PT Dobu ʔuwe rattan
PT Minaveha ue rattan
PT Tawala kuwe rattan
PT Saliba kuwe vine type, used for tying sago leafs to roof of bushmaterial houses
MM Bola hue Calamus hollrungii
MM Nakanai hue a thorny rattan, Calamus sp.
MM Lavongai ue rattan
SES Bugotu ɣue Calamus hollrungii, Calamus stipitatus and Calamus vestitus’ (W. McClatchey, pers. comm.)
SES Gela ɣue rattan
SES ’Are’are uwe a liana
SES Arosi ʔue rattan
SES Sa’a ue rattan cane’ (ACD)

6.3.2. Dendrocnide and Laportea spp., nettle trees, TP salat, filas, B nanggalat (Urticaceae)

Dendrocnide and Laportea species are nettles, i.e. plants with stinging hairs, often grouped together in the literature as ‘nettles’ or ‘nettle trees’. The hairs remain in the skin, enabling the toxin in them to spread. Those mentioned below range from the 40-metre canopy tree Dendrocnide excelsa to the small stinging herb Laportea interrupta. They are included here as shrubs simply because this is where a majority of their tokens belong.

Until 1965 the members of both genera were considered to belong to the genus Laportea, but, as a footnote by the translator, E.E. Henty in Peekel (1984: 151) explains, the genus Laportea was revised by Chew (1965), removing woody species from it and placing them in a new genus Dendrocnide. At the same time the genus Fleurya was abandoned and its species transferred to Laportea. Laportea are monoecious herbs whilst Dendrocnide are dioecious shrubs or trees.34 In both genera the fruit is dry and has a single seed.

Species of Laportea and Dendrocnide are often not distinguished by the glosses in the cognate sets below, but this at least in part reflects the usage of the terms. Wheatley (1992: 240) reports that in Vanuatu languages generally there is a single term for all Dendrocnide species, despite the fact that in Vanuatu they vary in size from the shrubby Dendrocnide latifolia, which occasionally grows to 10 m but is usually smaller, to the 25 m canopy tree Dendrocnide moroides.

Dendrocnide species are much better described in the literature than Laportea species, presumably because the latter are simply regarded as nuisance weeds. Species mentioned in the glosses below are, roughly from largest to smallest:

  • Dendrocnide excelsa (syn. L. gigas), the giant stinging tree, is a tree of the forest canopy, up to 40 m tall, with dull green heart-shaped or round leaves covered with stinging hairs.35
  • Dendrocnide harveyi (syn. Dendrocnide milnei, Laportea harveyi, L. milnei), a tree up to 20m tall, apparently found only in Fiji and western Polynesia.
  • Dendrocnide latifolia, a small shrubby tree, occasionally with a straight bole and reaching 10 m, but usually smaller. It is common in secondary forest. It has a serious sting, the pain of which lasts for days (Wheatley 1992:238, Kwa’ioloa & Burt 2001: 154, Scales n.d.).
  • Dendrocnide sessiliflora (syn. Laportea sessiliflora), a shrub or small tree 3-10 m high with a mild sting (Peekel 1984: 153).
  • Laportea interrupta (syn. Fleurya interrupta, Urtica interrupta) resembles a small European stinging nettle with a serious sting, and is cultivated and eaten by the Tolai of the Gazelle Peninsula of New Britain (Peekel 1984: 157).

Figure 7.11: Left Hornstedtia lycostoma: A, plant, about 3 m high, with suckers and inflorescences; B, inflorescences; C, longitudinal section oflarge fruit. Right Calamus hollrungii, rattan: A, leaf base and spikes; B, inflorescence; C, fruit; D, portion of leaf tip with barbs.

Apparently none of the trees is used for timber, but Ann Chowning (pers. comm.) reports that on New Britain Nakanai and Meramera speakers dry Laportea bark over a fire and use it for roofs and walls. Wheatley (1992: 240) comments that Dendrocnide moroides (syn. Laportea moroides), a canopy tree of 25 m found in central Vanuatu, is considered useless because the wood is very soft and rots quickly. The leaves and bark of a number of species have medicinal uses. The leaves of a Dendrocnide species were used medicinally on Manam Island. They were boiled in water or with grated coconut to cure constipation or general seediness (Wedgwood 1934: 286–287). Peekel (1984: 151) notes that the finely chopped leaves of Dendrocnide longifolia (a shrub or small tree, 3-5m high) were mixed in to dogs’ food to make them hunt more keenly. Dendrocnide latifolia was used in much the same way on Mwotlap, except that here the recipients were warriors: they were given a soup made from the leaves to render them quicker tempered and stronger in battle (Wheatley 1992: 238).

Several forms below reflect PAn *lateŋ with a prefixed syllable: POc *ja-latoŋ, PAdm *la-latoŋ, *ña-latoŋ, PNCV *ga-latoŋ. The existence of the various prefixed forms indicates that unprefixed *latoŋ was also inherited into POc, and this is borne out by a single unprefixed reflex below: Kove lato. POc *jalatoŋ, the most widely reflected form, must also have been inherited, as non-Oceanic reflexes occur. Apart from a few languages in the extreme north of Vanuatu - Vures, Mwesen, Dorig, Merlav - which reflect the prefix *ja- regularly (François 2004b), most known reflexes in Vanuatu, listed separately below, reflect *ga- ‘tree’.

PAn *lateŋ stinging nettle tree, Laportea harveyi’ (Blust 1972b)
PMP *zalateŋ Laportea and Dendrocnide spp.’ (Dempwolff 193 8)
POc *[ja]latoŋ Laportea and Dendrocnide spp.’ (Milke 1961: *salatoŋ; Ross 1989b)
PAdm *lalato, *ñalato Laportea and Dendrocnide spp.’ (Blust 1996b)
Adm Lou lalat Laportea and Dendrocnide spp.’ (Blust 1996b)
Adm Wuvulu lalaʔo Laportea and Dendrocnide spp.’ (Blust 1996b)
Adm Kele lulat Laportea and Dendrocnide spp.’ (Blust 1996b)
Adm Lenkau lalatr Laportea and Dendrocnide spp.’ (Blust 1996b)
Adm Seimat nalat Laportea and Dendrocnide spp.’ (Blust 1996b)
Adm Bipi ñalak Laportea and Dendrocnide spp.’ (Blust 1996b)
Adm Loniu ñalat Laportea and Dendrocnide spp.’ (Blust 1996b)
Adm Leipon ñilet Laportea and Dendrocnide spp.’ (Blust 1996b)
Adm Drehet nolok Laportea and Dendrocnide spp.’ (Blust 1996b)
PWOc *[ja]latoŋ nettle tree, Dendrocnide sp., perhaps Dendrocnide warburgii
NNG Kove lato Dendrocnide excelsa’ (Chowning 2001: 83)
NNG Takia dalat nettle tree
NNG Manam zalato tree sp. which causes itching, cooked and eaten after maternity, used as a medicine
NNG Sissano talat nettle plant; poison oak tree
MM Nduke zilatoŋo Laportea interrupta and Laportea ruderalis
MM Marovo zilatoŋo Laportea interrupta and Laportea ruderalis
MM Roviana jilatoŋo k.o. nettle
PEOc *[ja]lato nettle tree, Dendrocnide sp.
SES Gela (haŋa)lato Dendrocnide sp.’ (W. McClatchey, pers. comm.)
SES Sa’a (nunu)lao nettle tree
SES Ulawa (dū)lao nettle tree
SES Arosi darao k.o. nettle
NCV Vurës silat Dendrocnide sp.’ (François 2004b)
NCV Mwesen salat Dendrocnide sp.’ (François 2004b)
NCV Dorig (o)slat Dendrocnide sp.’ (François 2004b)
NCV Merlav ne-silat Dendrocnide sp.’ (François 2004b)
SV Sye n-elyat Dendrocnide sp.
Fij Wayan salato Dendrocnide harveyi’ (Geraghty 2004: 83)
Fij Bauan salato Dendrocnide harveyi’ (Geraghty 2004: 83)
Pn Samoan salato Dendrocnide harveyi’ (Whistler 2000: 196)
PNCV *ga-lato nettle tree’ (Clark 1996)
NCV Mwotlap na-hlat Dendrocnide spp.
NCV Mota kalato nettle tree
NCV Ambae kalato Dendrocnide latifolia
NCV Nokuku elat nettle tree
NCV Araki kalaro Dendrocnide sp.
NCV Tamambo (vu)kalato nettle tree
NCV Raga galato Dendrocnide latifolia
NCV Uripiv gelat nettle trree
NCV Labo na-ŋgalate stinging nettle
NCV Lonwolwol gela[r,t] stinging leaf bush

Clark (1996) plausibly suggests that PNCV *kara reflects POc *kaRat ‘bite’, and the final -t of Big Nambas n-harat appears to support this etymology.

POc *kara(t) a small stinging plant, perhaps Laportea interrupta’ (Chowning 2001: 83)
NNG Kove gala a stinging plant related to lato (Dendrocnide excelsa)?
MM Tolai kara stinging nettle spp., Dendrocnide sessiliflora, Laportea interrupta
NCV Tape nə-xārət stinging nettle
NCV Big Nambas n-harət stinging nettle
NCV Port Sandwich na-xer stinging nettle
NCV Paamese a-ai devil nettle (Dendrocnide sp.)
NCV Lewo ke nettle
NCV Namakir kar nettle tree
NCV Nguna na-kara nettle tree

6.3.3. Lygodium spp. (Lygodiaceae)

Ferns of the genus Lygodium are climbing vines which wind around tree trunks. Peekel (1984: 27, 30) writes that Lygodium circinnatum (syn. L. dichotomum, Lygodium flexuosum) grows 3-6m long, Lygodium trifurcatum, more delicate than Lygodium circinnatum, grows 3-5m high, and Lygodium scandens (syn. L. microphyllum) 1-3m long.

All are apparently used as binding material. The thicker Lygodium circinnatum and, among the Roviana, Lygodium trifurcatum are used for binding outrigger booms. The more delicate Lygodium scandens is used in weaving, for tying bundles and as circlets for the arm or leg (Waterhouse 1949, Arentz et al. 1989: 93).

PMP *qaRsam fern sp.’ (ACD)
POc *qasam fern used for tying and binding, Lygodium circinnatum’ (ACD; Chowning 2001: 83)
MM Nakanai hara Lygodium circinnatum
MM East Kara kasom Lygodium circinnatum
MM Patpatar sam Lygodium circinnatum
MM Patpatar sam-sam Lygodium scandens
MM Tolai em Lygodium circinnatum
MM Tolai em-ien Lygodium scandens
MM Tangga āsem the Lygodium creeper
MM Nehan heham Lygodium circinnatum
MM Petats aisam Lygodium circinnatum
MM Tinputz asam Lygodium circinnatum’ (Blackwood 1935)
MM Mono-Alu asama a creeper species’ (Record 1945)
MM Marovo amasa creeping coastal fern, Lygodium sp. (metathesis)
MM Roviana asama a trailing fern, Lygodium trifurcatum
SES Gela aha species of creeper used for stringe
SES Tolo asa vine sp. used to bind canoes and weave baskets
SES Kwara’ae sata Lygodium scandens’ (Whitmore 1966)

6.3.4. Merremia spp. (Convolvulaceae)

Merremia peltata (syn. Operculina peltata, Convulvulus peltatus, Ipomoea peltata) is a woody liana found throughout the rain forest, but it is particularly abundant in disturbed forest areas. The vine has the thickness of a human arm and grows 15-50 m high, with white-yellowish funnel-shaped flowers that resemble Morning Glory (Peekel 1984:467, Pawley & Sayaba 2003, W. McClatchey, pers. comm.).

According to Peekel Merremia peltata vines are used as binding material in circumstances where the fastening does not need to be durable. Hviding (2005: 124) reports from Marovo that older thicker vines contain a milky sap that is good for stopping blood flow in an emergency.

The Meso-Melanesian forms below other than Tolai valearu reflect POc *paliaRa, but reflexes of *paliaRua are found in New Ireland, southern Vanuatu and Fiji, and I take this to have been the POc form.

POc *paliaRua a vine, Merremia peltata
MM Nakanai valiala a vine, Ipomoea sp.’ (Floyd 1954)
MM East Kara viliai Merremia peltata
MM Madak leale Merremia peltata
MM Patpatar haliara Merremia peltata
MM Tolai valear[a,u] Merremia peltata
MM Ramoaaina waliara a creeping plant
SV Sye (nos-i)vilyau Merremia peltata
Fij Wayan veliawa Merremia peltata36

6.3.5. Asplenium nidus, bird’s nest fern (Aspleniaceae)

The bird’s nest fern, Asplenium nidus, usually grows as an epiphyte on the trunks or branches of trees in the rain forest or mangrove swamp. It has large simple fronds visually similar to banana leaves, growing to 50-150cm long and 10–20cm broad. The fronds are light green, often crinkled, with a black midrib. The fronds grow in clusters and roll back as they turn brown, creating a massive leaf nest where they are attached to the tree. It is an ideal understorey plant, as it thrives in warm, humid habitats in partial or full shade (Peekel 1984: 17, Hviding 2005: 124).

The POc term was *pʷete.

POc *pʷete bird’s nest fern, Asplenium nidus
MM Patpatar pate Asplenium nidus
MM Tolai pete Asplenium nidus
NCV Mota puɣet Asplenium nidus
NCV Dorig bıt Asplenium nidus’ (François 2004b)
NCV Lakon puıt Asplenium nidus’ (François 2004b)

7. The forest floor

Because it receives little light, the forest floor is often almost bare of plants that depend for their existence on photosynthesis. The lower parts of trees and the debris of fallen trunks and branches, however, provide a home for fungi of various kinds.

The generic term for mushrooms and fleshy fungi was POc *taliŋa (Ch 3, §4. 7), formally identical to *taliŋa ‘ear’ and presumably reflecting a perception that some fungi resemble the human ear. Blust (2000) points out that more detailed descriptions specify the referent of *taliŋa reflexes as jelly fungus, which do not have the umbrella-like shape of a mushroom but ‘sprout directly from the trunks of dead trees as a collection of folded tissues which may appear cup-like or ear-like’. Their names include reference to ‘ear’ in a number of cultures around the world. In Austronesian languages, including some in Oceania, the name also specifies the ear’s owner. Since they grow on tree bark, they are sometimes named ‘tree ear’. Whether one should therefore reconstruct POc *taliŋa qi kayu ‘tree fungus’ (lit. ‘ear of tree’) is a little debatable. This may well have been a POc locution, as Blust also notes its occurrence in non-Oceanic Austronesian languages, but it may also have been coined more than once in the history of Austronesian languages. Most of the terms below are from Blust (2000).

Adm Mondropolon can-n-i kei (cane-n ‘her/his ear’)
SV Anejom̃ in-ticŋa-nɣai mushroom (arboreal)’ (cf. in-ɣai ‘tree’)
Pn Māori tariŋa rākau

Some jelly fungi are named ‘rat ear’, apparently because of their shape:

NCV Paamese raliŋe-n asu kind of fungus which grows on dry wood
Mic Marshallese lɔcilŋi-n kicṛik toadstool, Auricularia ampla, and other ear-like Basidiomycetes (fungi)
Pn Rarotongan tariŋa kiore fungus sp. which grows on decaying trees
Pn Tuamotuan tariŋa kiore
Pn Māori taliŋa ʔimoa

A number of Oceanic terms translate as ‘ghost ear’. Blust suggests that this reflects their spiritual significance, associated with the hallucinogenic properties of some fungi and with their sudden appearance after a thunder storm - which also accounts for the ‘thunder ear’ terms below. The distribution of ‘ghost ear’ terms perhaps justifies the reconstruction of POc *taliŋa qi qanitu, literally ‘ear of spirit of dead’, but the term for ‘spirit of dead’ varies across languages.

Adm Seimat taxiŋ i paxi (lit. ‘ear of ancestral spirit’)
Mic Ponapean saleŋ en eni
Mic Mortlockese sᴂliŋa-n anu mushroom’ (lit. ‘ear of ghost’)
Mic Puluwatese hᴂliŋᴂ-n hoomᴂ tree fungus, mushroom’ (hoomᴂ ‘bad ghost of dead’)
Mic Chuukese seniŋe-r soomᴂ
Fij Rotuman faliaŋ ne ʔatua toadstool or fungus
Fij Bauan daliŋa ni kalou
Pn Tikopia tariŋa ŋa atua

The data below support a PMic reconstruction.

PMic *taliŋa ni para fungus growing on tree trunks’ (lit. ‘ear of thunder’)
Mic Kiribati taniŋa ni pa mushroom-like fungus growing on tree trunks, Myxomycetes: slime fungus’ (ba ‘thunder’)
Mic Woleaian taliŋe-ɾi-pac̣ mushroom’ (pac̣ ‘thunder’)
Mic Satawalese saliŋa-ni-pac̣ kind of toadstool

In our data sources terms for different kinds of fungus tend to be very few, and only one reconstruction is offered, based on work by French-Wright (1983).

PWOc *kokoi mushroom sp.’ (French-Wright 1983)
NNG Kove koko edible mushroom sp.’ (A. Chowning, pers. comm.)
PT Bwaidoga kokoio mushroom or toadstool sp.
MM Petats koko mushroom or toadstool

Notes